304 
Psyche 
[December 
species of Panchlora has all the phallomeres characteristic of most 
male blaberid genitalia. Five groups are erected for species of 
Panchlora , based on the number of phallomeres which are missing. 
The genital hooks (R2) are absent in Panchlora (from Central and/ 
or South America), Achroblatta , Biolleya, and Anchoblatta. 
The mating behavior of Panchlora differs from most other Blat- 
taria, and it is discussed in relation to the loss of the genital hook. 
Acknowledgments 
I thank the following for the loan of museum specimens: Dr. 
David Ragge (British Museum), Dr. Ashley Gurney (U. S. Na- 
tional Museum), Dr. Karl Princis (Lund), Dr. S. L. Tuxen 
(Copenhagen), and Dr. Frank Fisk. I am grateful to Mr. Samuel 
Cohen for taking the photographs. 
References 
Barth, R. Jr. 
1968. The mating behavior of Gromphadorhina portentosa (Schaum) 
(Blattaria, Blaberoidea, Blaberidae, Oxyhaloinae) : an anomalous 
pattern for a cockroach. Psyche 75: 124-131. 
Gupta, P. D. 
1947. On copulation and insemination in the cockroach Periplaneta 
americana (Linn.). Proc. Nat. Inst. Sci. India 13: 65-71. 
Khalifa, A. 
1950. Spermatophore production in Blattella germanica L. Orthoptera: 
Blattidae). Proc. Roy. Entomol. Soc. London 25(A): 53-61. 
McKittrick, F. A. 
1964. Evolutionary studies of cockroaches. Cornell Univ. Agr. Exp. Sta. 
Mem. No. 389, 197 pp. 
Princis, K. 
1960. Zur systematik der Blattarien. Eos 36: 427-449. 
1963. Orthopterorum Catalogus. Subordo Polyphagoidea : Fam. Homoe- 
ogamiidae, Euthyrrhaphidae, Latindiidae, Anacompsidae, Atti- 
colidae, Attaphilidae. Subordo Blaberoidea: Fam. Blaberidae. 
s’-Gravenhage, pp. 76-172. 
Roth, L. M. 
1969. The evolution of male tergal glands in the Blattaria. Ann. En- 
tomol. Soc. Amer. 62: 176-208. 
1970a. The male genitalia of Blattaria. III. Blaberidae: Zetoborinae. 
Psyche 77: 217-236. 
1970b. The male genitalia of Blattaria. IV. Blaberidae: Blaberinae. 
Psyche 77: 308-342. 
1971. The male genitalia of Blattaria. VI. Blaberidae: Oxyhaloinae. 
Psyche 78: 84-106. 
Roth, L. M. and H. B. Hartman. 
1967. Sound production and its evolutionary significance in the Blat- 
taria. Ann. Entomol. Soc. Amer. 60: 740-752. 
