212 
Psyche 
[Aug.-Oct. 
The selerite just behind the plate vs of Fig. 19 probably 
represents a modified coxite or protopodite of a limb (of the 
eighth uromere) in which the outer branch or exopodite was 
lost and the endopodite or inner branch became modified to 
form the ventral valve vv. The greatly elongated coxite or 
protopodite of the ninth uromere apparently formed the dorsal 
valve dv whose external branch or exopodite (represented by 
the stylus in immature roaches) becomes lost in the adult, 
while the inner branch, or endopodite, becomes modified to form 
the inner valve iv. This is a very complicated idea of the 
origin of the parts. A simpler view (though one which is less 
in accord with the facts of comparative anatomy) is that the 
ventral valves vv are merely outgrowths of the eighth uromere, 
while the dorsal valves dv are outgrowths of the ninth uromere, 
and the inner valvulse iv are outgrowths of the tenth uromere 
which become secondarily associated with the ninth segment. 
Miall and Denny state that “In the cockroach the ovipositor is 
used to grasp the egg capsule, while it is being formed, filled with 
eggs, and hardened: and the notched edge is the imprint of the 
inner posterior gonapophyses, made while the egg capsule is still 
soft. The shape of the parts in the male and female indicates 
that the ovipositor is passive in copulation, and is then raised to 
allow access to the spermatheca.” A richly branched valviglan- 
dula or valve gland ( vgl of Fig. 17), which is probably an ac- 
cessory colleterial gland, pours out its secretion through the 
valvipore vpo (Fig. 20) located between the bases of the inner 
valvulse iv. 
The ninth abdominal segment is the genital segment par 
excellence in the male, and it may be referred to as the andromere. 
As shown in Fig. 21, its pleurite pi is mueh enlarged and extends 
downward to the ninth sternite or hypandrium ha. The hy- 
pandrium ha of Fig. 21 consists of a basal portion or hypandrium 
proper, labelled eha and a posterior sex formed by the union of 
the coxites, or stylus-bearing rudimentary abdominal limbs (ves- 
tigial protopodites) which unite with a portion of the sternum 
to form the area labelled sex in Figs. 21 and 24, while the styli 
(vestigial exopodites) remain distinct to form the peg-like ap- 
pendages stl of the ninth sternite in the male. Normally the basal 
