82 
Psyche 
[June 
somewhat nearer than the Blattids are to the Orthopteroids, in 
the character of their maxillae. 
The character of the labium of Gryllohlatta (Fig. 1) clearly 
shows that this insect is Orthopterous or Orthopteroid, since it 
is only in the Orthoptera that I have found a separate and dis- 
tinct, well-chitinized mentum of the type shown in Fig. 1 , mn. 
The labium of Gryllohlatta lacks the transverse gular plate 
characteristic of all Dermaptera; and the type of labium ex- 
hibited by the Orthopterous insects might be derived more 
readily from Isopteroid or Blattoid precursors. 
It is rather surprising that the maxillae and labium of the 
Phasmids which I have examined are not as similar to these 
structures in Gryllohlatta as the maxillae and labium of the Em- 
biids are. This, however, is doubtless due to the fact that my 
material is not as suitable as it might be to indicate the real 
relationships involved, and there are certain features of the 
submental region of the Phasmid labium, for example, that 
indicate a much closer relationship to the Orthoptera should be 
expected in the Phasmids than in the Embiids. 
The mandible of Gryllohlatta (Fig. 3) is very like that of such 
Orthoptera as Gryllotalpa, and resembles the type occurring in 
the Embiids quite closely. The mandible of Gryllohlatta is not 
as much like that of the Phasmids and Dermaptera as one would 
expect, but it resembles the Dermapteran type as much as any. 
The • resemblance to the Isopteran or Blattid type is not very 
marked, so that the evidence of the mandibles it not of great 
phylogenetic value. 
The antenna of Gryllohlatta (Fig. 4) is remarkably like that 
of Emhia not only in the number of the segments composing it, 
but also in the relative sizes of the individual segments. The 
antenna of Gryllohlatta is also very like that of such Phasmids as 
Timema and this type of antenna was apparently inherited from 
a common Protorthopteroid source. Among the true Orthop- 
tera, the type of antenna found in the Acridids (Locustids) and 
their allies approaches the Grylloblattid type more closely than 
is the case with the antennse of the Tettigonioids and Grylloids, 
which is rather surprising in view of the fact that Gryllohlatta is 
more closely related to the Tettigonioid and Grylloid Orthoptera 
