12 
Psyche 
[March 
eastern New South Wales and southern Victoria; in these areas some 
species occur not only in rain forest (including south temperate rain 
forest) but also in wet sclerophyll forest and good savannah woodland. 
However only two groups of the genus reach Tasmania and only one 
group (two related, primarily allopatric species) occurs in rain forest 
there. 
Trichosternus is a genus of 25 or more species confined to eastern 
Australia, except that one species is isolated in southwestern Australia 
(Darlington 1953, p. 94). The genus’ northern limit is between 
Daintree and Cooktown. It occurs (several species, some very local- 
ized) throughout the main rain forest system of North Queensland, 
where it is apparently confined to rain forest. It is well represented 
also in the subtropical rain forest system of South Queensland and 
northern New South Wales, and in this area some species occur in 
savannah woodland as well as in tropical-type rain forest, and some 
have entered south temperate rain forest on the Dorrigo-Ebor plateau 
and the Mt. Royal Range. The southern limit of the genus is some- 
where in east-central New South Wales, probably not far north of 
Sydney. 
The northern limit of Leiradira (or of the group of genera that 
includes Leiradira) is between Daintree and Cooktown. This genus 
too occurs in much of the main tropical rain forest system of North 
Queensland, being represented there by several distinct species each 
more or less localized, but the genus may be absent in the southern 
extension of the main tropical rain forest system south of the Atherton 
Tableland. It is represented also by several species in the subtropical 
rain forests of South Queensland etc. Its southern limit is apparently 
on the lower, eastern edge of the Dorrigo plateau. It is confined to 
eastern Australia. It is wholly or chiefly a rain forest genus in all 
parts of its range. 
The three preceding genera are all Pterostichini. All their species 
are flightless geophiles. Additional flightless geophile pterostichines 
are localized in all the different rain forest areas of Australia from 
Cape York to Tasmania. Examples are Mecynognathus in the tip-of- 
peninsular forests; Paranurus in the mid-peninsular forests; Loxo- 
genius and undescribed genera in the main tropical rain forest system ; 
Nursus s. s.j Liopasa, Ceratoferonia Zeodera, and Notolestes in the 
subtropical rain forest system ; Loxodactylus in the wet forests of 
southern Victoria; and Rhabdotus ia those of Tasmania. (It should 
be added that Australia possesses many winged pterostichines as well 
as these and other flightless genera.) 
