i6 
Psyche 
[March 
is world-wide in distribution. It includes both flying and flightless 
genera, but the flying genera usually live beside standing or running 
water and are not forest-living geophiles. Flightless geophile Trechini 
are numerous both north and south of the tropics. In Tasmania they 
are numerous in south temperate rain forest and hardly enter other 
habitats at low altitudes, although some occur in open country above 
timber line, on cold mountain tops. Flightless Trechini are less numer- 
ous but still widely scattered in wet forests and on mountain tops in 
southern Victoria, including the Otway Ranges (Moore i960), east 
nearly to the New South Wales border and north to Mt. Kosciusko 
in southern New South Wales, and endemic species perhaps represent- 
ing one original flightless stock of spotted " Trechus ” are isolated on 
the Mount Royal Range, the Dorrigo-Ebor plateau, and the Mc- 
Pherson Range on the Queensland border. 6 
The tribe Migadopini (Jeannel 1938; Darlington i960, p. 663) 
is confined to the southern hemisphere, with different genera localized 
in Tasmania and southeastern Australia, New Zealand, and the south- 
ern tip of South America, etc. Two flightless genera of the tribe occur 
in Tasmanian rain forest: Calyptogonia is confined to Tasmania; 
Stichonotus extends to the mainland, but only to the Otway Ranges. 
A third Australian genus of the tribe is known from a single specimen 
collected long ago near Kiama south of Sydney, and a fourth genus 
occurs still farther north, in subtropical forest on the low (c. 2,000 
ft.) Comboyne plateau at about 3 1 0 35' S. This last genus, Decogmus, 
differs from all other Migadopini in being winged. 
Finally, the flightless tribe Agonicini is confined to Tasmania and 
southeastern Australia (Moore i960). There are two genera. One 
is widely distributed in Tasmania and occurs also in the mountains 
of southern Victoria east of Melbourne (B. P. Moore, in letter). 
The other is confined to the mainland, including the Otway Ranges 
and the “Victorian Alps,” north to Mt. Kosciusko. Agonicines live 
on the ground in rain forest, and sometimes in open snow gum woods 
on mountains. 
Although there are other Carabidae in Tasmanian rain forests 
(especially various Pterostichini and Licinini) the four tribes just 
discussed make up a large part, and zoogeographically the most im- 
portant part, of the flightless wet forest Tasmanian carabid fauna. It 
will be seen from details given above that all four tribes occur both in 
6 A second “Trechus”, diemensis Bates, extends from Tasmania and south- 
eastern Australia north to the McPherson Range, but this species is winged 
or dimorphic. 
