PSYCHE 
Vol. 68 December, 1961 No. 4 
AUSTRALIAN CARABID BEETLES VII. 
TRICHOS TERN US, ESPECIALLY THE 
TROPICAL SPECIES 
By P. J. Darlington, Jr. 
Museum of Comparative Zoology, Cambridge, Mass. 
This is one of a series of papers describing new Carabidae of zoo- 
geographic importance referred to in my account (1961b) of transi- 
tion of Australian wet forest carabid faunas. Some other papers of the 
series, including a list of my localities, are referred to below (p. 130). 
The present paper is concerned with Trichosternus , especially with 
the comparatively little known tropical species. Types of new species 
are placed, at least for the time being, in the Museum of Comparative 
Zoology; paratypes, with C. S. I. R. O. at Canberra and usually in 
the Queensland Museum. Proportions given in the descriptions are 
calculated from actual measurements. 
Items of geographic or evolutionary interest derived from Tricho- 
sternus are its total distribution ; the occurrence of two very distinct 
(relict or primitive?) species, fax and montorum , high on Mt. Bartle 
Frere (one reaching Mt. Bellenden Ker too) ; the distribution of T . 
f rater and its apparent allies ( suhvirens and simplicipes of South 
Queensland and nudipes to fisheri in the tropics) , which form a group 
of mostly allopatric forms in which the male front tarsi have apparent- 
ly been simplified in two different stocks and in which double invasion 
or hybridization may have occurred in a very limited area on part of 
the Atherton Tableland (pp. 122-125). Another case of hybridiza- 
tion (of eungella and mixtus ) may have occurred on the Eungella 
Range (p. 127). See map for distribution of species on and north 
of the Atherton Tableland. 
I cannot give an exclusive definition of Trichosternus. Characters 
used by Sloane (1894 etc.) and Tschitscherine (1902) fail among 
recently discovered species. Compared with Nurus, Trichosternus is 
usually smaller, more lightly built, with S front tarsi usually dilated 
and with 3 segments squamulose below, while in Nurus the cf front 
tarsi are usually simple, rarely (e.g. in N. atlas Cast.) slightly dilated 
and with 2 segments squamulose. However, exceptional Trichosternus 
