1961] 
Darlington — Australian Carabid Beetles 
ii5 
resemble Nurus in having cf tarsi simple ( T. subvirens Chd., sim- 
plicipes SL, nudipes Dari., soror Dari.) or with only 2 segments 
squamulose ( setosiceps SL). Compared with N otonomus, Trichos- 
ternus is usually larger, often with prosternal process setose (rarely 
in N otonomus) , often with alternate intervals of elytra raised, and 
often without distinct 10th intervals at outer edges of elytra, but all 
these characters fail in some species. A generic revision of the larger 
Australian pterostichines is much needed. The Australian genera 
should be compared with the New Zealand ones (Britton 1940) and 
phylogenies should be worked out if possible, and the old types, many 
of them in Europe, should be re-examined. However, I cannot under- 
take all this now. 
The known range of Trichosternus is the eastern edge of Australia 
from the base of the Cape York peninsula (south of Cooktown) to 
central New South Wales (perhaps a little north of Sydney), with 
one species isolated in southwestern Australia (Darlington 1953) and 
another on New Caledonia. All the tropical Australian species (ex- 
cept cordatus ) live in rain forest, but some more-southern species occur 
in savannah woodland, and some enter or are confined to south tem- 
perate rain forest on the Dorrigo-Ebor and Mt. Royal plateaus. 
Before considering the tropical Trichosternus, I give the following 
tentative key to species south of the tropics, as a basis for comparison 
( cf . Sloane’s key, 1899, pp. 567-569, and Tschitscherine’s, 1902, pp. 
Explanation of Plate 7 
Known distribution of Trichosternus in tropical Queensland. The finely 
dotted line is the approximate eastern edge of high land (Atherton Table- 
land etc.). Arrows indicate occurrence southward, at increasing distances 
beyond the limits of the map, of the species indicated. The species are num- 
bered in the order in which they are treated in the text. No. 1, on Mt. 
B(artle) F(rere), is Trichosternus fax ; 2, obscuripennis , which extends north 
beyond the limits of the map nearly to Cooktown; 3, montorum ; 4, nudipes, 
on the Mt. Spec plateau ; 5, soror ; 6, f rater, which occurs north to Mt. Lewis 
and south beyond the limits of the map on the Kirrama Range; 7, mutatus ; 
8, fisheri; 9, eungella, and 10, mixtus, both on the Eungella Range; 11, corda- 
tus, at the southern edge of the tropics; 12, spec, on the Mt. Spec plateau; 
13, setosiceps , which is widely distributed on the south-central Atherton Table- 
land and occurs also south of the limits of the map on the Kirrama Range ; 
and 14, kirrama, on the Kirrama Range. Nos. 1, 2, 3, 13, 14 are very distinct, 
phylogenetically isolated species. Nos. 4, 5, 6, 7, 8 are apparently interrelated, 
chiefly allopatric members of the frater group (see text). Nos. 9, 10, both on 
the Eungella Range, though structurally distinct, may be related to each other 
and may hybridize (see text). No. 11 represents a primarily south temperate 
rather than tropical stock. No. 12, on the Mt. Spec plateau north of Towns- 
ville, is apparently related to another south temperate (New South Wales) 
species. 
