1961] 
Darlington — Australian Carabid Beetles 
12 3 
transverse strigulation in baso-lateral impressions. Elytra slightly 
more than 1/4 wider than prothorax, subparallel, slightly narrowed 
anteriorly; margin forming obtuse (nearly right), slightly dentate 
angles at humeri ; dorsal striae formed by fine vaguely punctulate im- 
pressed lines in low spaces between intervals; latter subequal on disc, 
slightly raised but not costate, except 7th strongly raised and costate 
basally, 10th interval indicated posteriorly; each 3rd interval 1- or 
2-punctate posteriorly, position of punctures variable. Mesosternum 
anteriorly and prosternal process (sometimes inconspicuously) setose. 
Male front tarsi moderately dilated, 3 segments squamulose below; 
cf with 1, $ 2 setae each side apex last ventral segment. Length 
20-23 ; width 6. 5-8.0 mm. 
Holotype cf (M. C. Z. Type No. 30,358) and 73 paratypes all 
from Kirrama Range, inland from Cardwell, North Queensland, 
2000-3000 ft. altitude, Dec. 1957 and Feb. 1958, collected by the 
Darlingtons, in rain forest. Also the following specimens, not types: 
12, mountains above (SW of) Atherton, Atherton Tableland, 3000- 
4000 ft., Dec. 1957 and Feb. 1958; 1, near Black Mt., c. 20 miles 
north of Kuranda, probably near 2000 ft. altitude, May 1958; and 
3, Mt. Lewis, near Mossman, probably near 3000 ft., Dec. 1957; all 
collected by the Darlingtons, in rain forest. These localities are all 
on the Dividing Range system, which has probably been the principal 
path of north-south dispersal of wet forest Carabidae in North 
Queensland. 
T. frater may represent the ancestral stock, with dilated, squamu- 
lose male tarsi, from which nudipes of the Mt. Spec plateau, subvirens 
and simplicipes of South Queensland, and also (independently) soror 
of the Atherton Tableland etc. have been derived. These 5 species 
(including frater itself) are all allopatric and are generally similar 
in form and technical characters except for the differences in the cf 
tarsi. The two following new species may be local derivatives of the 
same stock with cf tarsi still dilated and squamulose. See map for 
distribution of tropical species of the frater group. 
Trichosternus mutatus new~ species 
Small; form about average for genus but rather convex; black, 
sometimes with faint purplish reflections; moderately shining, elytra 
slightly duller. Head 3/4 to 4/5 width prothorax; antennae rather 
short, scarcely extending beyond base of prothorax; eyes moderate, 
genae c. wide as eyes, rounded and narrowed to neck; 2 supra-ocular 
setae each side. Prothorax subcordate, 1/4 (db) wider than long at 
middle; base equal to or slightly narrower than apex; sides broadly 
