i8 
Psyche 
[March 
venation of the holotype is misinterpreted in the original figure 
(Guthorl, 1939, fig. 1), and the type specimen, which was deposited 
in a private collection, is not accessible for study. Hence, this species 
is referred to the Megasecoptera, family uncertain. 
Orthocosta lithomantidoides Waterlot, 1934 (Westphalian C, 
Belgium) is based upon a small wing fragment. Because of the 
shape of the posterior margin and the irregular cross venation, with 
incompletely developed intercalated sectors, I believe it belongs to 
the Megasecoptera, though of uncertain family position. Orthocosta 
splendens Bolton, 1912, on the other hand has a loose “palaeodicty- 
opterous” type of reticulation and is referred here to the Palaeodicty- 
optera, uncertain family position. 
Palaeopalara gracilis Handlirsch, 1904, and Antracopalara falci- 
pennis Handlirsch, 1904 (Westphalian C, Belgium) are small wing 
fragments with megasecopterous wing shape. Both species are here 
referred to the Megasecoptera, families uncertain. 
Since it seems clear that the most primitive Megasecoptera merge 
fully in the wing venation pattern with the Palaeodictyoptera, we 
must accept the fact that some little-known species cannot be identi- 
fied to either order with certainty. 
The following is a summary of my present concept of the relation- 
ships of the species mentioned above : 
Order Megasecoptera 
Family Dictyoneurellidae Kukalova-Peck (herein described) 
Dictyoneurella perfecta Laurentiaux, 1949 
Family Frankenholziidae 
Frankenholzia culmani Guthorl, 1962 
Family Mecynopteridae 
Mecynoptera splendida Handlirsch, 1904 
Family uncertain 
Saarodictyurn abnorrnis Guthorl, 1939 
? genus lithomantidoides Waterlot, 1934 
P dlaeopalara gracilis Handlirsch, 1904 
Anthracopalara falcipennis Handlirsch, 1904 
? genus tuber culata Bolton, 1921 
Order Palaeodictyoptera 
Family uncertain 
Orthocosta splendens Bolton, 1912 
Order uncertain 
Merlenbachia grimaldi Waterlot, 1934 
