56 
Psyche 
[March 
sternite as in Fig. 6, the dorsal flange of the stalk flaired out into 
membrane at the middle. Elytra covering all of the first four ab- 
dominal tergites, the fifth tergite partially exposed. Folding pattern 
of hind wing similar to that illustrated for Anthobium (— Eusphal - 
erum ) sorbi by Forbes (1926: fig. 33), the first transverse fold being 
a hinge in which the costal margin is turned about 90°. 
Tergites of abdominal segments 2 and 3 sclerotized to some degree, 
those of fourth and following segments more strongly so. Tergites 
4 and 5 each with a pair of patches of short, pale, mesally-directed 
setae, the patches together covering most of the surface of the tergite. 
Sternite 8 with complex anterior modification as in fig. 7. Inter- 
segmental membranes of abdomen with rectangular plates arranged 
in a brick-wall-like pattern. 
Male genital segment and aedeagus as illustrated by Jeannel 
(1962: figs. 10, 11), except that both figures are ventral rather than 
dorsal views as stated. Pleurites of genital segment narrowly con- 
nected anterior to tergite. Parameres of aedeagus in repose in ab- 
domen facing dorsally. Female external genitalia as in figs. 8-10. 
Discussion 
Jeannel considered Glypholoma to be closely allied to the silphid 
tribe Pterolomini. He differentiated it from that tribe by the struc- 
ture of the maxillary palpus, with a very short third segment, and 
by what he considered a very unusual aedeagus. Scheerpeltz placed 
the genus in the Omaliinae and considered it closely related to 
Lathrimaeum (— Anthobium of current usage). He separated it 
from Anthobium by the absence of a nuchal sulcus and by the longer 
and differently sculptured elytra. 
In considering the placement of the two long-controversial genera 
Brathinus and Camioleum , Hammond (1971) has reviewed and 
augmented the characters of the Omaliinae that are useful in dis- 
tinguishing them from other Staphylinoidea. Glypholoma has all 11 
of the mentioned features. The three characters that will immediately 
exclude Glypholoma from association with Pterolomini and allied 
tribes of Silphidae are : 1 ) the hinging of the costal margin of the 
hind wing during folding, whereas the costal margin is folded back 
on itself in primitive Silphidae as illustrated for Necrophilus hydro- 
philoides by Forbes (1922: fig. 27) ; 2) the anterior modification of 
the eighth sternite, whereas Silphidae have at most a simple median 
projection; and 3) the structure of the aedeagus with a large lightly 
sclerotized basal bulb, short median lobe and small median foramen, 
