352 
Psyche 
[September-December 
Each body segment with 12 skin folds; folds neither diverging 
nor fusing along lateral side above the legs. The skin papillae with 
round bases but of varying sizes and shapes; lateral papillae with 
more elongate and straight-sided tip, tips shorter and more rounded 
on dorsal papillae; one to three but usually two small papillae be- 
tween major papillae. 
Mandible (Figs. 5, 6) outer blade with long thin main tooth and 
a short thin accessory tooth ; inner blade with long, thin main tooth 
and two thin shorter accessory teeth. No small sawlike teeth below a 
diastema on outer or inner blade. 
Type locality description and observations. The type locality 
cave is 1500 feet long, at 1700 feet elevation, 0.6 air miles SE of 
the village of Pedro River, in northeastern Clarendon Parish. The 
cave is a large diameter, relatively simple, moist tunnel with large 
deposits of calcium carbonate dripstone and bat guano, and an air 
temperature of 2i.5°C. All specimens were found near the middle 
of the cave in the “Three Ways” section (see map in Fincham and 
Ashton, 1967). One was found in the open on wet decomposed 
guano before, and the other three specimens in the chamber after, 
the low crawlway leading to Three Ways. In 1974, one specimen 
was sighted while it was partially exposed, feeding on an immature, 
blind Nelipophygus roach ( Blatellidae) , with most of its body hid- 
den in a small hole at the side of a ceiling pocket. The second 1974 
specimen was found hidden under a loose clay-guano flake formed 
by the drying of a formerly flooded section of the cave floor. Four 
people searched this chamber thoroughly for an hour in 1974 for 
more specimens but found none. A rich invertebrate fauna of po- 
tential prey exists in the cave. When touched, all specimens ejected 
a viscous, sticky fluid for 2-3 cm from their oral papillae. 
Cave adaptation. The eyeless and depigmented conditions of the 
species are usually associated with troglobitic (cave-evolved) species, 
but are not limited to such species. These characters are also en- 
countered in forest litter-evolved species, and many litter inhabitants 
in the past were certainly ancestral to presently troglobitic species 
(Barr, 1968). The legs and antennae of Speleoperipatus may be 
somewhat elongated (Fig. 8) as a specialization for cave life, but 
measurements (in life) of the appendages of this and epigean species 
are not available for comparison. 
The general region of Jamaica from which Speleoperipatus comes 
is known to have several species of seemingly troglobitic invertebrates 
(personal observations), all of which have been derived from litter- 
inhabiting ancestors. But collecting in forest litter in Jamaica is 
