1988] 
Cornell, Stamp, & Bowers — Hemileuca lucina 
53 
Table 4. Relationship between instars for total distance traveled by marked 
individuals in groups. Spearman rank coefficients (r s ) and probability (P) at a = 
0.05 are indicated. 
Group 
Instar 
n 
r s 
P 
A 
1-2 
42 
0.227 
0.15 
2-3 
34 
0.218 
0.22 
3-4 
27 
0.308 
0.12 
4-5 
20 
0.097 
0.69 
B 
1-2 
36 
0.156 
0.37 
2-3 
36 
-0.176 
0.31 
3-4 
32 
0.668 
0.001* 
4-5 
32 
0.403 
0.02* 
C 
1-2 
40 
0.177 
0.28 
2-3 
34 
0.313 
0.07 
3-4 
33 
0.447 
0.01* 
4-5 
27 
0.418 
0.03* 
D 
1-2 
- 
- 
- 
2-3 
25 
0.369 
0.07 
3-4 
23 
0.446 
0.03* 
4-5 
21 
0.303 
0.19 
♦Indicates statistical significance 
between larval weight and distance traveled for Instars II-V, and 
exhibited a significant correlation between distance traveled between 
instars for two such combinations. In contrast, Group A had only 
one instar in which rankings were stable among trials and one instar 
in which distance traveled was correlated with weight. 
In H. lucina, transient polyethism occurred at several levels: 1) 
Over the course of a trial, larval ranks did not change; thus, leaders 
during a single foraging bout (represented here by a single trial) 
remained at the forefront. 2) In some cases, ranks of individual 
larvae were constant from one trial to the next within an instar. The 
instar in which this was most prevalent was the third (for 3 or 4 
groups). However, in other cases, there was no consistency in larval 
ranking from trial to trial. Thus, in some groups the same indi- 
viduals led a series of foraging bouts, whereas in others, the leaders 
varied from bout to bout. In some instances, a significant positive 
correlation occurred between larval weight and distance traveled, 
but in other cases, there was no correlation. Thus, within a foraging 
group, the largest larvae were sometimes the leaders, sometimes 
randomly distributed within the traveling group, but were never 
consistently the slowest (i.e. none of the negative correlation coeffi- 
