1988] 
Binder — Aggression in Heterotermes 
129 
Table 1. The evasion of soldier aggression by workers during all intercolony 
pairings of workers and soldiers. 
Arena size (cm 2 ) 
Worker evasion 
no. a 
% a 
1.5 
4 
20 
3.0 
3 
15 
4.5 
4 
20 
6.0 
9 
45 
Total 
20 
25 
a Each value based on 20 pairs of termites 
Table 2. Comparison of worker behavior and survival during all intercolony 
pairings of soldiers and workers. 
Worker response 
no. 
% 
active 
Initial behavior 
44/80 
55 
Within group survival 
17/44 
39 
Total survival 
17/80 
21 
passive 
Initial behavior 
36/80 
45 
Within group survival 
3/36 
8 
Total survival 
3/80 
4 
Table 3. The aggressive response of soldiers to immobilized workers and the 
aggressive response of workers to immobilized soldiers. 3 
Immobilized workers 
Immobilized soldiers 
Abdomen 
exposed 
Head 
exposed 
Abdomen 
exposed 
Head 
exposed 
Aggressive response 
on first encounter 
3 b 
5 
3 
7 
Aggressive response 
after 3 encounters 
3 
5 
10 
9 
a Aggressive responses followed encounters of individuals of different colonies and 
resulted in a snap of the mandibles by the soldier or a worker attack 
b Each column represents the actual number of individuals responding of 10 pairs of 
termites 
