1988] 
Carpenter — Gayellini 
227 
species, recognition of Paragayella is but another example of the 
current needless fragmentation of vespid generic classification, 
which I have decried elsewhere (Carpenter and Cumming, 1985; 
Carpenter, 1987, 1988). A fully sequenced cladistic classification 
(Wiley, 1979) is possible with a single genus. I am therefore 
synonymizing Paragayella with Paramasaris. 
Gayella 
The monophyly of the genus is shown by the pointed clypeus 
(Figs. 14-17; not similar to that of Stenogastrinae and Polistinae), 
the temples projecting somewhat and the emarginate and bispinose 
last metasomal tergum (Figs. 50-52). In other Masarinae and 
Euparagiinae the clypeus is truncate or emarginate, the temples do 
not project and the last visible metasomal tergum is neither 
emarginate nor spined. Several characters of the male genitalia are 
also synapomorphies. The digitus is enlarged relative to Paramasaris 
(cf Figs. 56 and 57-63), the cuspis is tuberculate basally (Figs. 
57-61, 63), and the paramere has an enlarged lobe ( dorsal to the 
spine, Fig. 62). The combination of features of the male genitalia is 
unique in Vespidae. Finally, the globose shape of the first 
metasomal tergum may be apomorphic, but this is variable within 
the genus (Figs. 43-45). 
Within the genus, two monophyletic species groups may be 
recognized, which allows a classification that is phyletically se- 
quenced (Wiley, 1979). These are the eumenoides group, for 
eumenoides and araucana, and the mutilloides group, inlcuding 
reedi, patagonica, luispenai and mutilloides. 
Eumenoides group 
The monophyly of the eumenoides group is established by the 
tegula, which lacks the short posterior lobe found in Paramasaris 
and the mutilloides group (cf. Figs. 47 and 48). This feature is 
approached in reedi (Fig. 49), diminishing its strength. The 
projection from the cuspis is tuberculate and apical relative to that 
view of metasomal terga I and II. 42. P. fuscipennis, 13X. Lateral view of meta- 
somal tergum II. 43-44. Dorsal view of S metasomal tergum I, 10X. 43. Gayella 
araucana. 44. G. eumenoides. 45. G. luispenai 9, 10X. Posterodorsal view of 
propodeum and metasomal tergum I. ap: anterior pronotal carina ;/• pronotal fovea; 
It: longitudinal carina of metasomal tergum II; mg: propodeal median groove; 
pp: posterior pronotal carina; TI: metasomal tergum I; 77/: metasomal tergum II. 
