230 
Psyche 
[Vol. 95 
derived features, however. The preoccipital carina is evanescent and 
separated from the postocular carina by more than an ocellar 
diameter in species of this group, whereas both are well developed 
and closely approximated in the eumenoides group and Euparagii- 
nae. The femur is punctate in the mutilloides group; it is smooth in 
other gayellines. The spines defining the emargination of the last 
visible metasomal tergum are narrow and elongate in the mutil- 
loides group (Fig. 50), whereas they are broader and shorter in the 
eumenoides group (Figs. 51-52). Since both states uniquely charac- 
terize monophyletic groups, the polarity cannot be clearly inferred. 
However, the state of the metasomal spines in the mutilloides group 
is a more extreme development, and is here suggested as relatively 
apomorphic. 
Within the group, reedi is the sister-group to the remaining three 
species. I have not discovered any clear autapomorphies of this 
species. Synapomorphies uniting patagonica, mutilloides and luis- 
penai include greater development of the long black hairs on the 
metasoma (extending over the disc of tergum II, Fig. 53), and an 
elongate malar space (length 1 / 2 to greater than the width of the 
interantennal distance, Fig. 14; shorter than this in other gayellines, 
Figs. 15-17). The postocular carina tends to be more effaced (as in 
Fig. 20), but traces appear to be present in some specimens. Meta- 
somal sternum II in the male and to some extent also the female is 
bordered posterolaterally with blunt ridges (Figs. 53-54), however 
these are variably developed in patagonica, and reedi approaches 
this condition. Among these three species, the features I have polar- 
ized are autapomorphies.' Hence, the relationships are at present 
unresolved (Fig. 2). Autapomorphies of the species are: for mutil- 
loides the very long malar space (Fig. 14), the acroglossal buttons 
more elongate and the glossa more deeply bifid than other Gayella, 
and the cuspis tubercle quite blunt (Fig. 61); for luispenai the male 
metasomal sternum II projections elongate (Fig. 54); and for pata- 
gonica the female propodeal median groove narrowed before 
broadening dorsally (Fig. 55; smoothly narrowed in other Gayella, 
Fig. 45). 
Identification Keys 
Genera 
1. Pronotum with two transverse carinae (Figs. 29-31); clypeus 
emarginate to truncate (Figs. 11-13); last tergum neither 
