1972] 
Miller & Kurczewski — Entomognathus 
73 
weight ratios, expressed as decimals, for E. memorialis, Crossocerus 
maculiclypeus (Fox), and Lindenius columbianus errans (Fox) 
were 0.77, 0.22, and 0.07, respectively. This point should not be 
stressed, however, because the weight of the female wasp may vary 
depending upon her physical condition (see Iwata, 1942). We 
caught a female of Cerceris fumipennis Say (Sphecidae) at Auburn, 
N. Y., weighed, released, and recaptured her 12 days later, and 
found that her body weight had decreased from 95 to 74 mg, or 
22 % ! 
Many fully-provisioned cells of E. memorialis lacked an egg, 
larva, or parasites, and we believe this may also be related to the 
physiological state of the female. Because they were excavated 
late in the nesting season, such cells were possibly made by females 
which had already exhausted their egg supplies. 
The rather small mean number of prey per cell recorded for 
E. memorialis (5.1) is undoubtedly related to the relatively large 
mean sizes of the beetles. E . brevis constructs a cell of similar size 
but stores, on the average, considerably more (17.8), usually smaller 
beetles per cell. There is, in fact, no overlap in the ranges of the 
number of prey per cell for brevis (14-24) and memorialis (3-9). 
Several other differences in nesting behavior are evident between 
brevis and memorialis , particularly in nest structure and dimen- 
sions. Projecting turrets have been noted for nests of the former 
but not for nests of the latter. Series of cells in tandem are common 
in nests of memorialis but have not been observed in nests of brevis . 
The maximum number of cells per nest in memorialis (20) is twice 
as high as that reported for brevis (10). The burrows of the former 
are longer than those of the latter, and the nests are more compli- 
cated. Although both species place the egg transversely across the 
mesosternum of the beetle, the cephalic end is attached mesad to the 
base of the procoxa in brevis (see Fig. II in Grandi, 1927), whereas 
in memorialis it is fastened laterad of the base of the procoxa 
(Fig- 3)- Furthermore, the caudal end of the egg extends beyond 
the side of the beetle in brevis while in memorialis it does not. 
Several noteworthy similarities in the nesting behaviors of brevis 
and memorialis may be applicable for the entire genus. Both 
species paralyze their prey, probably by stinging, and enter their 
nests in flight, holding the beetle with the middle legs. The burrow 
storage of prey observed in memorialis is similar to that reported 
by Maneval (1937) for brevis. The burrow-searching adults of 
brevis observed by Adlerz (1912) may play a role similar to that 
