1984] 
Aiello — Genus Adelpha 
39 
6 Morrell (1954) gives a fascinating account of larval behavior in 
several Malayan genera of the tribes Limenitidini (Athyma, 
Moduza, Pandita ), and 7 Neptini (Lasippa, Neptis, Phaedyma). 
Except for Neptis leucoporos, the larvae, of the butterflies he 
observed, expose the midrib, or a secondary leaf vein, by feeding 
around it (just as Adelpha does); young larvae rest out on these 
slender supports. Like Adelpha also, Athyma, Moduza, and 
Pandita accumulate their feces to form a mass at the base of the 
support vein; Neptini do not. 
Judging from Morrell’s illustrations, the larva of Moduza 
resembles a combination of Adelpha cocala and basiloides, es- 
pecially in having short thick scoli on A2; its pupa is remarkably like 
A. cocala or leucophthalma, but with slightly stalked head horns; 
the host plants are mainly Rubiaceae (Mussaenda, Nauclea, 
Timonias, Uncaria, Wendlandia), with one record on Oleaceae 
(Olea). 
The larvae of Athyma kanwa feeds on Uncaria, and is described 
by Morrell (1960) as “brown with delicate green branched spines, 
and resembles a growth of moss on a decaying patch of leaf;” the 
pupa (illustrated in Morrell, 1954) is silver with long head horns and 
to me looks for all the world like a pupa of Adelpha celerio. 
In form, Athyma nefte larvae (Morrell, 1954) are reminiscent of a 
very plump Adelpha iphicla larva, and have been reported on 
Glochidion (Euphorbiaceae) and Mussaenda (Rubiaceae); the pupa 
is “golden brown in colour, with two dorsal plates of brilliant 
metallic gold,” and from what can be determined from its picture, it 
looks like the pupa of Adelpha iphicla or cytherea, but with an 
exagerated A2 projection, and like those species it appears to have 
short pointed head horns. 
Pandita sinope is described by Morrell (1954) as having a larva 
similar to that of Athyma nefte in color and general appearance; the 
illustration of the pupa is also similar to A. nefte, but has a less 
6 According to Corbet and Pendlebury (1978), five of the seven names used by 
Morrell (1954) should be amended: his Neptis columella = Phaedyma columella; 
Neptis heliodore = Lasippa tiga; Neptis nata = Neptis leucoporos; Parathyma 
kanawa = Athyma kanwa; Parathyma nefte = Athyma nefte. His Moduza procris 
and Pandita sinope remain unchanged. 
7 Neptini is defined by Eliot (1969) to include Aldania, Lasippa, Neptis, Pantoporia, 
and Phaedyma. 
