1984] 
Aiello — Genus Adelpha 
3 
the Archives of the Museum of Comparative Zoology, Harvard 
University. About half of this manuscript is in a nearly illegible 
hand; half is typewritten. The typed portion includes part of an 
introduction, notes on each species, a key (based upon wing pattern) 
to species and species groups, and descriptions of four new species. 
Included also are more than 70 genitalic illustrations (inside of right 
valve) by Howarth. Forbes had analyzed these latter and was in the 
process of constructing a key to genitalia. His two approaches, of 
wing pattern and genitalia, yielded different species groupings, and 
it is not clear whether he favored one of these, or intended to use a 
combination of the two in his final classification. 
Of the immature stages of Adelpha, very little has been said, and 
what has been said has been largely ignored for the simple reason 
that this new information appears to confuse matters rather than 
clarify them. 1 Forbes (uncompleted manuscript) had read Moss’s 
(1933) paper on Adelpha larvae and pupae, and had examined 
several pupal skins of Adelpha and Old World Limenitini in the 
collections of the British Museum (Natural History) when he 
commented, “The larvae and pupae are highly varied and unless a 
high percent are misdetermined, show characters wholly incongru- 
ent with adult structures and patterns.” 
The fact that, within Adelpha, the study of adult characters 
results in species-alignments different from those obtained by 
consideration of the immature stages and/or genitalia, suggests that 
at least one set of characters is unreliable or perhaps even 
deliberately deceptive. 
Immature Stages 
Due to taxonomic confusion within Adelpha, it is impossible to 
know how many species actually are represented by the 34 or so life 
history accounts published for this genus. My estimate is that at 
least 24 species are illustrated (including illustrations in this paper) 
as final instar larva, or pupa, or both, many of these by more than 
one author. Most accounts, whether illustrated or not, include a 
foodplant record. 
'This author has located only four publications (Moss, 1933; Muller, 1886; Young, 
1974; Comstock & Vasquez, 1960) which figure any immature stages of Adelpha; 
several publications present descriptions only. 
