310 
Psyche 
[Vol. 9! 
In a series of papers, Thornton and co-workers (Thornton and 
Smithers 1974, 1977, 1978, Thornton and New 1977a, b, Thornton, 
Wong, and Smithers 1977) have described numerous species of 
Philotarsidae from Australia, New Zealand, New Guinea, New Cale- 
donia, and surrounding islands. Following these works, Thornton 
(1981) presented a classification of the Philotarsidae. 
In an earlier work, Lee and Thornton (1967) described numerous 
species of Pseudocaeciliidae from southeastern Asia and the Islands 
of the South Pacific. If one compares the figures in this paper with 
figures for the Philotarsid genera Zelandopsocus and Austropsocus 
in the papers cited above, one sees numerous points of marked 
similarity. 
Methods 
The observations of notable similarities between two Philotarsid 
genera and several Pseudocaeciliid genera led me to make a compar- 
ison of as many of the genera of both families as the literature and 
material at hand allow (Table 1). Comparisons were made of 11 
characters. All of those chosen hold constant (i.e., in the same state) 
in more than one genus but fewer than all genera of the assemblage. 
Therefore, they may be expected to hold information about rela- 
tionships among these genera. Among characters which qualify in 
this way, the ones chosen are easily defined, and most of them have 
been described and/or figured in recent taxonomic works. Illustra- 
tions (Figs. 1-3) are included for characters which require them. 
Results and Discussion 
In eight of the characters, both Austropsocus and Zelandopsocus, 
with three tarsomeres, agree with the Pseudocaeciliid genera. In 
presence of only a single row of setae on the M-Cu stem in the 
forewing, Zelandopsocus agrees with the other Philotarsid genera, 
while Austropsocus agrees with the Pseudocaeciliid genera. In the 
lateral tyne of the lacinial tip, Zelandopsocus, from published fig- 
ures, appears to be intermediate between the other Philotarsids and 
the Pseudocaeciliids (but observation of Z. cumulus Thornton and 
New indicates closer proximity to Pseudocaeciliids), while Austrop- 
socus agrees with the Pseudocaeciliids. Bryopsocus agrees with the 
other Philotarsids in five characters, with the Pseudocaeciliids in 
