1984] 
Moekford — Psocoptera 
311 
two, is clearly intermediate in two, while two characters remain 
unknown in this genus. 
I conclude on the basis of these comparisons 1) that Austropsocus 
and Ze/andopsocus, find their closest affinities with the Pseudocae- 
ciliidae and should be placed there; 2) Bryopsocus is in several 
respects intermediate between these two families; it is unusual in 
several respects (see diagnosis, below) and apparently is best placed 
in a distinct family; 3) the forms remaining in Philotarsidae are the 
genera assigned to Subfamily Philotarsinae by Thornton (1981) 
(Thornton’s other Subfamily, Zelandopsocinae, consists of Ze/and- 
opsocus, Austropsocus, and Bryopsocus). 
It is necessary, then, to assign the genera placed in Philotarsidae 
by Thornton (1981) to three families: Philotarsidae Pearman, 
Bryopsocidae new family, and Pseudocaeciliidae Pearman. The 
three families may be diagnosed as follows: 
Philotarsidae Pearman (type Psocus picicornis Fabricius). Most 
“outer” and all “inner” sensilla of distal margin of labrum occupying 
a trough in margin, 4 on its outer and 5 on its inner wall; 2 most 
lateral sensilla on outer surface of labrum (total of 1 1 — “Type 1” of 
Table 1) (Fig. 1); lacinial tip with lateral tyne relatively broad; tarsi 
in adult 2- or 3-segmented; pretarsal claws each with a distinct 
preapical denticle and a slender pulvillus; M-Cu stem in forewing 
generally with one rank of setae; abdomen lacking eversible vesicles; 
aedaegus rounded apically; external parameres not protruding 
much beyond tip of aedeagus; hypandrium lacking paired lateral 
lobes; subgenital plate one-lobed distally; second valvula bearing a 
low, rounded lobe medio-distally. Included genera: Philotarsus 
Kolbe, Aaroniella Moekford, Broadheadia Moekford and Evans 
(regarded as a synomym of Aaroniella by Thornton), Haplophallus 
Thornton, Tarsophilus Moekford and Broadhead, Latrobiella 
Thornton. 
Bryopsocidae new family (type Austropsocus townsendi Smith- 
ers). Lacinial tip with lateral tyne somewhat less broad than in 
Philotarsidae; tarsi in adult 3-segmented; pretarsal claws each with a 
distinct preapical denticle; M-Cu stem in forewing with two ranks of 
setae; Cu2 in forewing with or without setae; hypandrium lacking 
paired lateral lobes but with a single low, rounded lobe on each side; 
aedeagus pointed apically; external parameres protruding well 
beyond tip of aedeagus; subgenital plate one-lobed distally; second 
