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Psyche 
[Vol. 94 
Table 4. The lower left portion of the matrix contains the probabilities that male 
and female regression lines in each comparison have the same slope. The upper right 
portion contains probabilities that male and female regression lines have the same 
intercept. Intercepts have been computed only for those cases where slopes are hom- 
ogenous. PW = pronotum width, PL = pronotum length, EW = elytra width, EL = 
elytra length, ED = elytra diagonal, FM = femora length. 
Morphological Character 
PW 
PL 
EW 
EL 
FM 
PL 
<0.001 
— 
0.252 
0.008 
0.628 
EW 
<0.001 
0.866 
— 
<0.001 
0.027 
EL 
<0.001 
0.146 
0.169 
- 
0.061 
FM 
<0.001 
0.417 
0.534 
0.226 
ED 
0.004 
0.002 
<0.001 
<0.001 
<0.001 
Some aspects of beetle shape changed abruptly with incresing size 
in some individuals and were impossible to quantify. For instance, 
the elytra surface near the humeral angle which is an evenly curved 
surface in small and intermediate males is a complexity twisted 
surface in large males. Additionally, an oblong perforation as 
large as 2.5 X 1.5 mm was present in approximately the same 
location in the elytra of many males (Fig. 4). The abdomen lies just 
central to the holes and apparently is never penetrated. Only males 
whose elytra length exceeded 8 mm were perforated (Fig. 6). The 
elytra of recently eclosed, large males (elytra length 8 mm) often 
lacked any perforation or occasionally had a single narrow slit in 
one side. Recently opened holes in young individuals sometimes had 
the appearance of having been punched inward from the dorsal side. 
The holes of older males were jagged in outline and widened into 
elipses. The shape of a few holes resembled a cross indicating that a 
planar object had been inserted and then twisted by 90 degrees. 
Sectioned elytra from young males showed clearly that the thickness 
of the elytra varies considerably and is thinnest in the area where 
holes normally occur. 
Interactions at the oviposition site 
Individual males, large enough to be easily distinguished from 
females, tended to reside on leaves toward the apex of M. umbellata 
shoots where ovipositing and guarding females were also common 
(Table 1). Activity increased noticeably between 1000 and 1200 hrs. 
