1987] 
Peeters & Crewe — Ophthalmopone berthoudi 
211 
relationships, attempts should be made to understand how it might 
have arisen as an effect of various attributes of the life histories. 
Acknowledgments 
We are grateful to William L. Brown Jr. for his information on 
the phylogenetic relationships among the Ponerinae, and to Barry 
Bolton for ant identifications. Peter Goodman provided logistic 
assistance in the field, and the Natal Parks Board gave permission to 
work in Mkuzi Reserve. Anthony Bannister took the photograph in 
Figure 1. We thank R. H. Crozier, W. L. Brown, and M. W. Moffett 
for comments on this manuscript. This study was funded by grants 
from the University of the Witwatersrand and the C.S.I.R., and the 
manuscript was prepared while CP was supported by the Australian 
Research Grants Scheme. 
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