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Psyche 
[Vol. 94 
some of the veins are raised areas or impressions depending on 
which face is viewed. Vague outlines of the head and sucking beak 
are indicated near the basal end of the antennae. 
Diagnosis. Brodioptera stricklani is closely related to B. cum- 
berlandensis Copeland 1957. Comparison of the wing of B. cumber- 
landensis (Carpenter, 1963) reveals that it more closely resembles 
the forewing of B. stricklani than the hindwing. The two may be 
separated on the basis of details of forking patterns of the veins. The 
shorter subcosta of B. stricklani joins the costa at about the mid- 
point of the wing while that of B. cumberlandensis joins the costa 
well beyond the midpoint. The radial sector, media, and cubitus all 
fork more distally in B. stricklani than in B. cumberlandensis. Both 
the fore and hind wings of B. stricklani have a relatively broader 
anal region than does B. cumberlandensis. Further, the MA in B. 
stricklani does not curve forward as sharply as in B. cumber- 
landensis. 
Etymology. This species is named in honor of Mr. Dave Strick- 
en of Provo, Utah, who donated the holotype specimen for this 
study. 
Discussion. The well-spread wings of B. stricklani give supple- 
mentary support to the concept that Megasecoptera were unable to 
fold their wings over the dorsum of the abdomen and, therefore, 
that Carpenter (1963) was correct in placing Brodioptera in Mega- 
secoptera. Further, the adequately preserved genitalia of this speci- 
men closely resemble those of extant Ephemeroptera (Edmunds, et 
al., 1976) and should provide valuable information for future phy- 
logenetic and evolutionary studies. It is interesting to note, as did 
Carpenter (1963), that such apparently apomorphic reduction in 
venation is present in an insect found so near the age (Namurian) in 
which the oldest winged insects have been discovered. 
Additional material. A second specimen consisting of genita- 
lia and cerci has been collected near the same locality as the holo- 
type. The specimen is considerably smaller than the holotype but 
has a similar genitalic structure. This specimen consists of both the 
part and counterpart that reveal the dorsum of the specimen 
including the dorsal attachment of the cerci to the abdomen. This 
second specimen is tentatively assigned to B. stricklani and is depos- 
ited as figured specimen #3243 in BYU paleontological collection. 
