1976] 
Aspey — Response Strategies of Schizocosa 
95 
ing in the female’s vicinity. Thus, the extensive agonistic behavioral 
repertoire may also confer a mating advantage to those males most 
successful in driving other males away from the female. 
Although this hypothesis has not yet been demonstrated experi- 
mentally, the males do exhibit stable, linear dominance-subordi- 
nance relations among themselves when tested in the laboratory 
under varying social, spatial, and population densities (Aspey, 
1975b, 1976b, c). Using a variety of analytical and descriptive tech- 
niques, each adult male S. crassipes can be classed as Dominant, 
Intermediate, or Subordinate in terms of the kinds and frequencies 
of agonistic behaviors exhibited during male-male encounters. 
Although the dominance classification is based on one animal’s 
responses to an opponent (i.e., inter-individual behaviors), this 
study demonstrates that strikingly different response strategies 
are also adopted by Dominant and Subordinate spiders when 
intra-individual behavior sequences are analyzed 
Methods 
Subjects 
The subjects were 40 adult males of the ground-dwelling brush- 
legged wolf spider Schizocosa crassipes (Walckenaer) (body 
length = 7-10 mm; carapace width = 3-4 mm; leg span = 27-30 mm). 
Molting to the adult occurs in late May, at which time the males 
develop conspicuous tufts of black hairs on the tibiae of the fore- 
legs, with smaller brushes on the patellae. The spiders were col- 
lected as immatures among leaf litter in early May at Stroud’s Run 
State Park, Athens, Ohio, U.S.A., and housed individually in 
visually-isolated, covered plastic containers (12.5 X 7.0 X 7.0 cm) 
until 1 wk after the final molt. Seven to ten days after the final molt 
each spider was marked with nontoxic enamel paint (“Pactra 
’namel,” Los Angeles, CA) on the dorsal surface of the cephalotho- 
rax and/or abdomen. No detrimental behavioral effects were ap- 
parent following paint application. 
All spiders were tested in four groups each of two, three, or five 
spiders matched into size categories of 0.5 mm increments based 
on adult carapace width (Hagstrum, 1971). Behavioral observa- 
tions were made in various sized rectangular glass terraria (90, 180, 
270, or 540 cm 2 floor space) having paper substrata strewn with 
