1976] 
Aspey — Response Strategies of Schizocosa 
101 
Intra-individual behavioral sequences are diagrammed separately 
for the 16 Dominant spiders (Figure 1) and 16 Subordinate spiders 
(Figure 2). Dominant spiders were characterized by those agonistic 
behaviors comprising Factor I (Approach/ Signal) and Factor II 
(Vigorous Pursuit) of Aspey’s (1976b) factor analytic classifica- 
tion. Subordinate spiders were characterized by Run, Retreat, and 
Vertical Extend, those behaviors comprising Factor III (Run/ Re- 
treat) of Aspey’s (1976b) factor analysis of the agonistic behavioral 
repertoire. A comparison of the original 20 behaviors displayed 
by Dominant and Subordinate spiders revealed that Dominant 
spiders exhibited a wider variety of behaviors linked together (15 
out of a possible 20) than did Subordinate spiders (10 out of 20). 
Thus, these results indicate that Dominant and Subordinate spiders 
can not only be reliably identified on the basis of responses made 
by conspecifics toward them during agonistic interactions (Aspey, 
1976b), but also by the response strategy exhibited during intra- 
individual behavior sequences. 
Discussion 
A transition probability matrix of agonistic behaviors exhibited 
during adult male-male interactions in S. crassipes indicated that 
a wider variety of responses followed any given behavior by a single 
spider during intra-individual sequences than during inter-indi- 
vidual encounters (Aspey, 1976b). However, examination of Table 
I revealed that some of the sequential responses appeared inconsis- 
tent (e.g., Run or Vibrate-Thrust following the performance of 
Horizontal Extend; Run or Front Approach following the peform- 
ance of Vibrate-Thrust). Horizontal Extend and Vibrate-Thrust 
are signal and active pursuit behaviors, routinely followed by some 
form of retreat during inter-individual encounters (Aspey, 1976b). 
However, if an animal exhibited Horizontal Extend or Vibrate- 
Thrust, seemingly aggressive behaviors, why would it suddenly 
retreat immediately after performing such a behavior? 
Further analysis examining each specific spider indicated that 
some spiders were consistently Dominant over all partners, while 
others were consistently Subordinate to every partner. For ex- 
ample, Dominant spiders not only performed Horizontal Extend 
or Vibrate-Thrust more frequently, but also continued to exhibit 
Vigorous Pursuit behaviors if an initial encounter with a conspecific 
