1976] 
Carpenter — Permian Insects 
349 
ferous, is only sparsely represented in Permian strata. Two species 
have been reported from the Elmo beds. 3 One of these, Dunbaria 
fasciipennis Tillyard, is a typical spilapterid. The other, Kansasia 
pulchra Tillyard, was originally described as a spilapterid (Till- 
yard, 1937) but has subsequently been transferred by Demoulin 
(1954) to the extinct order Archodonata. Although incompletely 
known, pulchra is almost certainly closely related to the species 
on which the order Archodonata was based (Martynov, 1931), but 
I do not believe that any of these insects are sufficiently distinctive 
to justify separation from the Palaeodictyoptera. Accordingly, I 
consider the Archodonata to be synonymous with the Palaeodic- 
tyoptera. 
Among the fossils which I collected at Elmo in 1927 is a speci- 
men that shows both palaeodictyopterous and megasecopterous 
traits. No additional specimens of this insect have subsequently 
been found, but I am now quite convinced that it is a representa- 
tive of a new family of Palaeodictyoptera. 
Family Elmoboriidae, new family 
Fore wing: slender, at least four times as long as wide. Costal 
margin nearly straight; SC long, terminating a short distance be- 
fore the wing apex; R1 close to and nearly parallel to SC, except 
distally; RS arising about !4 wing length from base, with several 
branches; M forking nearly at the same level as the origin of RS; 
MP forking almost immediately after its origin, with 2 or 3 
branches; CU forking near the base of the wing; CUA and CUP 
deeply forked; at least 2 anal veins. The costal margin of the wing 
is conspicuously serrate, the posterior margin less so. Hind wing 
and body unknown. 
This family differs from others previously described in the 
Palaeodictyoptera by the elongate shape of the wing, which is 
broadest distally, and by the proximity of the forking of M to the 
first fork of MP. The family is based on the genus Elmoboria, 
described below; the genus Oboria Kukalova (1960), from the 
Permian of Moravia, apparently belongs here also. 
3 A third species, Permoneura lameerei, was doubtfully placed in the Palaeodic- 
tyoptera (Carpenter, 1931) but as pointed out below (p. 373) the unique specimen 
on which the species was based does not show the diagnostic features of the order. 
