1978] 
Kugler — Pogonomyrmex mayri 
181 
species I was able to examine directly, the head is at best only 
slightly longer than wide in some species (Cl range of 4 Ephebo- 
myrmex species 89-97). The mesoscutum is long (mesoscutal L/WL 
33-50%), propodeum short (propodeal L/WL 26-37%) and the 
petiole has a distinct node. The legs are shorter than in mayri (hind 
femur L/WL 61-76%), and the front wing usually has a r-rn 
crossvein that joins the Rs at or distal to the end of the first cubital 
cell (see Cole, 1968: 25-26, plate 1 fig. 1). If the r-m vein is absent, 
the first cuboidal cell is open. 
Pogonomyrmex mayri may in fact deserve full generic status, but 
for the present it seems prudent not to create a monotypic genus 
before Pogonomyrmex is completely revised. The most recent 
revisions (Kusnezov, 1951; Cole, 1968) have been regional in scope 
and thus inadequate to address the question of whether the sub- 
genera Pogonomyrmex and Ephebomyrmex are really two distinct 
genera. Should such a split occur, mayri would no doubt be placed 
in the separate genus Forelomyrmex. 
Acknowledgements 
I thank Norman F. Johnson for taking the SEM photographs, 
Janet M. Hahn for helping prepare various stages of the manuscript, 
W. L. Brown, Jr. for his advice and constant support, and E. O. 
Wilson for the loan of specimens from the MCZ. This research was 
supported by the NSF grant DEB-22427 (W. L. Brown, Jr., 
principal investigator). 
References 
Brown, W. L. 
1953. Revisionary studies in the ant tribe Dacetini. Am. Midi. Nat. 50: 1-137. 
1975. Contributions toward a reclassification of the Formicidae. V. Ponerinae, 
tribes Platythyreini, Cerapachyini, Cylindromyrmecini, Acanthostichini, 
and Aenictogitini. Search, Agrculture, Cornell Univ. 5(1): 1-116. 
Cole, A. C. 
1968. Pogonomyrmex Harvester Ants, A Study of the Genus in North 
America. Univ. of Tennessee Press, Knoxville. 222 pp. 
Creighton, W. S. 
1952. Studies on Arizona ants (3), the habits of Pogonomyrmex huachucanus 
Wheeler and a description of the sexual castes. Psyche 59: 71-81. 
Forel, A. 
1899. Biologia Cent. -Am., Insecta, Hym. 3 (Formicidae), pp. 1-169 + 4 pi. 
