1978] 
Brartdao — Worker Caste of Formica perpilosa 
231 
Table 1. Relative frequencies of behavioral acts by the three worker categories of 
a single colony of Formica perpilosa. (N, total number of behavioral acts recorded 
for each caste). Approximate population of the nest: one nest queen, 150 minor 
workers, 270 media workers, 30 major workers, 50 eggs, 7 larvae and 5 pupae. 
Behavioral Acts 
A — Grooming 
1 — Autogrooming head, 1st pair of legs 
2 — Autogrooming 2nd pair of legs 
3 — Autogrooming abdomen, 3rd pair 
of legs 
4 — Allogrooming minor workers 
5 — Allogrooming media workers 
6 — Allogrooming major workers 
7 — Allogrooming nest queen 
B — Brood Care 
8 — Standing at the brood pile 
9 — Carry egg (or eggs in succession) 
10 — Lick egg (or eggs in succession) 
1 1 — Carry larva (or larvae in succession) 
12 — Feed larva 
13 — Lick larva (or larva in succession) 
14 — Carry pupa 
15 — Lick pupa 
16 — Assist eclosion to adult 
C — Regurgitation Behavior 
17 — Regurgitation with minors 
18 — Regurgitation with medias 
19— Regurgitation with majors 
20 — Donating to the Queen 
21 — Lay trophic egg 
22 — Feed queen trophic egg 
23 — Feed larva trophic egg 
24 — Carry infrabucal pellet 
25 — Feed on infrabucal pellet 
D — Working 
26 — Foraging outside the nest 
27 — Feed on diet 
28 — Feed on honey 
29 — Feed on cockroaches 
30 — Carry live nestmate 
31 — Carry dead nestmate 
32 — Drag the nest Queen 
33— Handle nest material 
34 — Carry debris 
35 — Lick nest wall 
36 — Excavating 
E — Other Behaviors 
37 — Antennal tipping 
38 — Jittering 
Total 
no. behavioral categories observed 
per caste 
minor 
media 
major 
workers 
workers 
workers 
N = 996 
N = 1679 
N = 134 
.167 
.131 
.224 
.060 
.062 
.052 
.023 
.022 
0 
.054 
.019 
0 
.043 
.095 
.052 
.007 
.012 
.030 
.011 
.049 
.022 
.066 
.077 
.134 
.033 
.055 
0 
.016 
.053 
0 
.009 
.024 
0 
0 
.004 
0 
.011 
.031 
0 
0 
0 
0 
0 
0 
0 
.002 
.024 
0 
.070 
.024 
.052 
.040 
.165 
.291 
.007 
.021 
.045 
0 
.009 
.008 
0 
0 
0 
0 
.001 
0 
0 
.001 
0 
.003 
.009 
.090 
.007 
.011 
0 
.082 
.021 
0 
.022 
.004 
0 
.029 
.025 
0 
.020 
.007 
0 
0 
.001 
0 
0 
.003 
0 
0 
.001 
0 
.018 
.005 
0 
.033 
.003 
0 
.059 
.034 
0 
.092 
.012 
0 
.001 
.002 
0 
.016 
0 
0 
1.0 
1.0 
1.0 
28 
34 
11 
