Colony l.a-C was queenless. 
Code Behavior 
Personal Behavior 
RE Rest 
SC Self-Groom 
MO Move inside Nesi 
{N=18) (N=t4) 
1519 1104 2078 
.2155 .2211 .1378 
.2892 .3298 .3584 
Brood Care 
IE Inspeci Egg 
CE Groom Egg 
CE Carry Egg 
LE Lay Egg 
AL Assist Egg-Lay 
ying 
GL Groom Larva 
CL Carry Larva 
RL Regurgitate w Larva 
FLD Feed Larva Solid 
ALE Assist Larval Eedysis 
IP Inspect Pupa 
CP Groom Pupa 
CP Carry Pupa 
AAE Assist Adult Eclosion 
.0058 
.0032 
.0088 
.0061 
.0061 
.0415 .03.36 0292 
,0461 03.36 0428 
.0058 .0071 0149 
.0084 0052 0061 
.22 
.06 
.07 
07 
.07 
0026 0006 
.0019 .0052 
0039 0039 
0006 0006 
0019 
.05 
ALWAIIogroom Worker 
BC Be Groomed 
ATQ Aniennaie Queen 
ALQ Allogroom Queen 
RQ Regurgitate w/Queen 
FQ Fight Queen 
0142 
0026 
0006 
.0013 
0048 
ATM Aniennate male 
ALMAIIogroom Male 
CM Carry Male 
RM Regurgitate w/ Male 
ALF Allogroom Alale Female 
Physical Nest Maintenance 
LN Lick Nest Walt 
LOT Look Outside Nest 
LEx Inspect Exuvium 
CEx Carry Exuvium 
EEx Eat Exuvium 
HM Handle Nest Material 
CR Carry Refuse 
FF Forage 
MOTMovc Outside 
IDr Inspect Prey 
CDr Carry Prev 
EDr Eat Prey 
FdN Feed Inside Nest 
FDO Feed Outsde Nest 
IFd Inspect Food 
DR Drink 
.07 
0045 
Social Organization in Lepioi 
