1983] 
Herhers — Social Organization in Leptothorax 
383 
coefficient, geometric distance between colonies failed to produce 
separate species clusters (Figure 6). Interestingly, queenright colo- 
nies of L. amhiguus clustered, as did monogynous colonies of L. 
longispinosus. However, an anomalous cluster was comprised of 
La-C and Ll-D, thereby reducing the significance of other clusters 
in the dendrogram. In sum, consideration of ethogram frequencies 
produced a dendrogram for which within-species variation swamped 
between-species variation. 
A final dendrogram was produced from comparison of time 
budgets over all colonies (Figure 7), based again on geometric dis- 
tance. This analysis was more discriminating than that based on 
ethograms (minimum similarity = 83.17). The dendrogram shows 
three longispinosus colonies clustered closely and three amhiguus 
clustered closely. The single aberrant entry was Ll-D, a polygynous 
lOOn 
q: 
< 
if) 
Ld 
> 
!< 
cd 
99 - 
98 - 
97 
La-A La-B La-C Ll-D Ll-C Ll-A Ll-B 
Figure 6. Dendrogram of similarity derived from geometric distance based on 
ethogram frequencies. 
