1983] 
Calahi, Traniello, and Werner — Pheidole 
411 
Mirknda, J. T. and S. B. Vinson. 
1981. Division of labor and specification of castes in the red imported fire ant 
So/cnop.sis invicia Buren. Anim. Behav. 29 : 410-420. 
OSTl R. G. F. AND E. O. Wll SON. 
1978. CaMc and Ecology in ihe Social Insects. Princeton University Press, 
Princeton, N.J. 
PoRTI R, S. D. AND C. D. JoRGHNSf N. 
1981. Foragers of the harvester ant, Pogonomynne.x owylieei: a disposable 
caste? Behav. Ecol. Sociobiol. 9; 247-256. 
Sin I V, T. D. 
1982. Adaptive significance of the age polyethism schedule in honeybee colo- 
nies. Behav. Ecol. Sociobiol. II: 287-293. 
Tranifi I (), J. F. A. 
1978. Caste in a primitive ant: absence of age polyethism in Anihlyopone. 
Science 202 : 770-772. 
Wll SON, E. O. 
1971. The Insect Societies. Belknap Press of Harvard University Cambridge, 
MA. 
1974. The soldier ant Camponotns (Colohopsis) fra.xinicola as a trophic caste. 
Psyche M 182-188. 
1975. Sociohiology: The New Synthesis. Belknap Press of Harvard University. 
1976a. Behavioral discretization and the number of castes in an ant species. 
Behav. Ecol. Sociobiol. 1: 141-154 
1976b. A social ethogram of the neotropical arboreal ant Zacryptocerus varians 
(Fr. Smith). Anim. Behav. 24 : 354-363 
1978. Division of labor in fire ants based on physical castes ( Hymenoptera: 
Formicidae:5'o/c/?D/M/.v). , 1 . Kansas Entom. Soc. 51 : 615-636 
1980a. Caste and division of labor in leaf-cutter ants ( Hymenoptera:Formici- 
d'de.Atta). 1. The overall pattern in A. se.xdens. Behav. Ecol. Sociobiol. 
7: 143-156. 
1980b. Caste and division of labor in leaf-cutter ants ( Hymenoptera:Formici- 
dae:/l//r7). 11. The ergonomic optimization of leaf cutting. Behav. Ecol. 
Sociobiol. 7: 157-165. 
Wll . SON, E. O. AND R. M. Faghn. 
1974. On the estimation of total behavioral repertories in ants. N. Y. Entomol. 
Soc. 82 : 106-112. 
Appendix I 
Hue and degree of pigmentation for the color (age) classes of the two physical 
castes of P. hortensis. 
Minors. 
Class /-White-yellow: head and thorax, pale white/ light grey; gaster, light 
grey; petiole and femur, pale white/ light yellow; tibia, grey. 
