132 Psyche [Sept.-Dee. 
praeterminal mark, and “trickling” distad across the terminal 
space (one, or a pair, on each side of the interneural fold). 
VUIb. Second , Third , Discoidal , and Lateral Macules. 
Second macules: if a Lycceides forewing is placed with its 
base towards the observer and its discal constellation is viewed 
from an imaginary horizontal line joining the opposite ends of 
the discales, the second macules , (R 3 ), R 4 , M 4 , M 2 , M 3 , and 
Cui 15 all of which have radiated from positions adjacent to 
cell RM, are seen to form a fairly regular rather weak arc, 
sloping somewhat sideways in relation to the rheniform 
macule RM, as if tipped by a slight apical pull (process 3). 
The twin macules Cu 2 and 1A lie outside the lower end of the 
arc, i.e.j lead an independent existence, having reached their 
present position (phylogenetically, from an enormously remote 
starting point in comparison to the starting points of the other 
macules) in result of a (process 2) cubito-anal stretch of the 
membrane (so conspicuously retained in some genera) that had 
occurred at some period in the evolution of the narrow and 
ovoid ancestral forewing (with a similarly shaped hindwing) 
prior to the comparatively recent generic and tribal apical de- 
velopment (process 3) which in a way has tended to repeat the 
initial growth and elongation of the ancestral wing (process 1). 
These stages of unequal growth and of subsequent compensa- 
tory readjustment may be compared to the already discussed 
evolutionary phenomena in the case of the uncus. 
It would be necessary to analyze a great number of ge- 
neric patterns (in the Plebejince alone striking variations on a 
P-shaped basis occur in Agriades , and a remarkable apicoid 
angle is formed by the macular constellation in Albulina ) in 
order to bring out certain features of the position of second 
macules in Lycceides , but this would transcend the scope of 
this paper. In selecting the three positions (1 proximal, 2 cen- 
tral, and 3 distal) given for this genus, stress has been laid on 
the progress of macule Cu 4 , but actually this may be combined, 
at these and intermediate stages, with shiftings on the part of 
the anterior series which may be removed from RM further 
than it is shown here. Fig. 2a shows the generic starting point 
of R 4 whose initial rather distal position (in regard to even such 
15 When no Roman number is appended to the symbol of the macule, the 
reference is to the second macule (e.g. Cui = Cui II). 
