Nest Defense Forager Nest 
Species site 1 type 2 type 3 spacing 4 Evidence 5 Source 
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acervorum distribution 1956a) 
Tetramorium A b,c?,e?,f? II o/+ Morisita’s index, tend to Baroni-Urbani (1969) 
caespitum overdispersion 
Tetramorium A,B b,c,e,f II o/+ Intra- and interspecifically Brian et al. (1966) 
caespitum territorial 
