Nest Defense Forager Nest 
Species site 1 type 2 type 3 spacing 4 Evidence 5 
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5 V/M, variance to mean ratio; !, our measurements or statistics based on published data; R, Clark and Evans’ nearest 
neighbor analysis, R = 1, random dispersion, R > 1, overdispersed, R < 1, clumped 
6 Mabelis (1979) studied this species in dunes; it is usually found in woodland. 
