372 
Psyche 
[Vol. 88 
them up and carrying them around until they eventually discard 
them, usually unharmed, at the garbage dump. It is conceivable that 
the carrying posture of Atemeles has evolved from a defensive 
“death feigning” behavior employed by less advanced ancestral 
species. 
Finally, Pella beetles do not have adoption glands associated with 
trichrome bristles. It is most likely, however, that the small clusters 
of glandular cells in the paratergites (for example in P. humeralis ) 
represent morphological precursors of the massively developed 
adoption glands in Atemeles and Lomechusa. 
Acknowledgements 
We would like to thank Hiltrud Engel for technical assistance, 
Dr. V. Puthz for identifying the beetles, Ruiko Pierce for translating 
the work of Yasumatsu from Japanese into English, and A1 Newton 
and Margaret Thayer for helping in disentangling the segmental 
morphology of Pella. This work was supported by grants from the 
Deutsche Forschungsgemeinschaft and National Science Founda- 
tion. 
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