68 
Psyche 
[March 
notauli obsolete; parapsidal furrows present but inconspicuous. Vena- 
tion nearly “complete/’ with the median abscissae of Rs (Rsf2 # 3) 
missing, so that the cubital cell is undivided (single). The first 
abscissa of M forks off from Cu basad of crossvein cu-a, as in the 
army ants. (This pattern of venation does not hold for T. pusillus, 
in which the veins are reduced and their relationships modified.) The 
hind wing entirely lacks an anal lobe, but has the large discal cell, 
usually with 2 or 3 stubs of apical abscissae corresponding to Rs, M 
and Cu; another small cell may occur at the base of the discal cell 
behind, or may be incorporated into the discal cell. The hamuli 
number 3, and usually arise from a small darkened sclerotic patch a 
little beyond the midlength of the costal margin. 
In other characters, gynes resemble workers. 
Male: (Based on T. rogenhoferi and T. clavicornis ) smaller and 
more slender than the corresponding gyne, but the difference is slight 
in the smaller species; dark brown to black in color; habitus typical 
of proponerine males. Eyes large and hairy, occupying nearly half of 
the sides of the head. Ocelli distinct. Scapes straight, of moderate 
length, usually equal to about the basal 3 or 4 flagellar segments; 
flagellum 12-segmented, the segments all longer than broad and 
increasing very slightly in thickness toward the apex. Mandibles well- 
developed, opposable or crossing at closure, dentition a variably 
reduced copy of that of the corresponding worker. Palpi segmented 
1, 2 (rogenhoferi) or 1, 1 (clavicornis). Clypeus with a large, 
swollen mid section and small sunken side pieces. 
Alitrunk with notauli developed only as the anterior arms of the 
“Y” and obsolete medially; parapsidal furrows present but inconspicu- 
ous. Wings as in gyne (see above). Legs slender, all three pairs with 
tarsal claws toothed. 
Petiole subclavate, i.e., with the peduncle rising gradually caudad 
toward nodal summit, which is rounded; ventral tooth or process 
present. Gaster with a slight constriction behind postpetiole; the 
latter segment has tergum and sternum firmly fused, but the next 
segment, abdominal IV, appears to have them connected only by thin 
cuticle or membrane. Genitalia only partly retractile, with parameres 
broadly rounded at apex; volsellae varying with the species (Figs. 7, 
8) ; aedeagal valves ordinary, serrate. Hypopygium with a long, more 
or less digitiform, upcurved, hairy, median process (ventral view, 
Fig. 9 )- 
Head longitudinally striate, rest of body predominantly smooth 
and shining, with fine scattered punctures. Pilosity fine, mostly erect 
and rather short, abundant and generally distributed. 
