A REVISION OF THE ANT TRIBE CARDIOCONDYLINI 
(HYMENOPTERA, FORMICIDAE) 
I. The Genus Prosopidris Wheeler 1 
By Jonathan Reiskind 
Biological Laboratories, Harvard University 
This is the first of a series of papers revising the tribe Cardio- 
condylini, a group of small myrm'icine ants (up to 3.0 mm in length), 
characterized by the possession of a pedunculate petiole, a wide 
postpetiolar dorsum, propodeal spines, and a general lack of pilosity. 
The tribe currently embraces the genera Cardiocondyla Emery 
(virtually cosmopolitan, mainly Old World), Xenometra Emery 
(West Indian) and Prosopidris (Papuasian). Xenometra includes a 
single poorly known species which is apparently parasitic on Cardio- 
condyla emeryi (Emery, 1917). Cardiocondyla and Prosopidris are 
free living, and form small inconspicuous colonies usually in the soil 
or in rotting logs. 
In this contribution Wheeler’s subgenus Cardiocondyla (Prosopi- 
dris) is elevated to full generic status. The worker and female types 
of its single previously known species, P. sima Wheeler (Philippines), 
are redescribed and a second species, P. papuana (New Guinea), is 
newly described. The papuana material includes the first known male 
of the genus, along with females and workers. The male is highly 
ergatoid, like those of some Cardiocondyla species. 
Measurements and Indices: In order to characterize properly cardio- 
condyline ants the following measurements and indices (with their 
abbreviations) will be used in this and succeeding papers: 
Head Length (HL) — Maximum length of head in frontal view 
from clypeal apex to posterior border of 
occiput. 
Head Width (HW) — Width of head measured in frontal view 
immediately behind the eyes. 
Scape Length (SL) — Maximum measurable length of scape, not 
including its articular condyle. 
Weber’s Length (WL) — Weber’s length of mesosoma ( = ali- 
trunk), measured in lateral view along a 
line connecting the place where the prono- 
tum joins the cervix and the apices of the 
Research supported by U.S. National Science Foundation Grant No. 
GB1634. 
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