48 
TRAUB & AUDY 
an adjacent microtibiala ; tarsus with a well-developed tarsala, a distal microtarsala, a subterminala and a 
parasubterminala and a pretarsala. Leg II, a genuala; two tibialae; a tarsala and proximal microtarsala, 
a pretarsala. Leg III, a genuala and a tibiala. 
Type Material. — Holotype and 5 paratypes from an unidentified bird, Mt Kinabalu, Tenompok, 
elev. 4500 ft., 1. viii. 1951 ( R . Traub). Holotype and paratype inU.S. National Museum (U.S.N.M. 2013); 
one paratype in collection of senior author, remainder in coll, of Colonial Office Medical Research Unit. 
Comment. — A series of Trombicula from crow-pheasant collected at Beaufort, N. Borneo, 
iii. 1952 (AL Nadchatram) superficially closely resembles T. vorca n. sp. As can be seen from 
the following standard measurement, the scutum is definitely and significantly smaller, as are 
the scutal-bristles : 
AW PW SB ASB PSB A-P AL AM PL DS PW PW PW PW 
Cox II SD ASB A-P 
Mean 69 80 28 32 12 35 50 52 64 52 1.2 1.2 2.2 2.35 
Range 2322124354 0.04 0.05 0.07 0.05 
The posterior eyes in the Beaufort specimens are fairly well developed and on an ocular 
plate with the very large anterior eyes. In the smaller scutal measurements the Beaufort 
series are close to T. }densipiliata , Womersley 1952, from the Celebes and the Solomons. 
Mr. Womersley has very kindly lent two specimens of his }densipiliata i but has been 
unable to locate T. nissani Dumbleton 1947. There is no doubt that T. densipiliata Walch, 
T. ? densipiliata, Womersley, T. nissani , T. vorca n. sp., and the form under study from 
Beaufort are all closely related. The results of further studies will be published later. 
A number of nymphs of the Beaufort species or variety were bred by Mr. Nadchatram. 
These nymphs, to be described later, are very distinctive, with exaggerated humeral and poste- 
rior dorsal body-setae. They are close to T. ( Neotrombicula ) canestrinii (Buffa, 1899), described 
from adults from soil in Italy. 
Trombicula (Leptotrombidium) baluensis n. sp. (figs. 25-33, pp. 58, 59). 
Diagnosis of Larva. — Unusual among chiggers of Southeast Asia in that the dorsal setae 
are very heavily barbed with thickened blade-like processes thus resembling species charac- 
teristic of Japan and Korea. Near T. intermedia Nagayo et al. (fig. 38-41, p. 60), and scutellaris 
Nagayo et al. (fig. 34-37, p. 60) of Japan, but immediately separable by virtue of the fact that in 
the new species the barbs of the dorsal bristles are less dense, far fewer in number per shaft 
and spaced further apart, i.e., the distance between barbs often 4-5 times their diameter at 
base instead of being almost contiguous ; the barbs are much longer than in the Japanese forms., 
i.e. twice the diameter of the shaft or greater, blade-like, the seta appearing subpectinate or 
with teeth like a rake instead of the barbs being subequal to the diameter of the shaft. Further 
distinguishable from T. (L). intermedia by the fine of SB lying anterior to PLs instead of behind 
it as in intermedia — the difference expressed by A-P of about 30 instead of 24, or the ratio 
PW/A-P being 77/30=2.6 not 78/24=3.2. T. baluenis is characterized by a relatively 
narrow AW — about 60, instead of 69-73 T. scutellaris and T. intermedia. 
Description of Larva. — Body: ovate in unengorged specimen, about 220x150(4. Ocular plate 
virtually contiguous with scutum, PLs in line with junction of anterior and posterior eyes. Gnathosome: 
Palpal formula N(/ow < g)/N/NN.B(ca. 7). Palpal claw apparently bifid, with lateral prong straight and 
dorsomedial prong smaller, often recurved under lateral. Galeal seta B {ca. 8, fairly long). Scutum: 
About 40 x 8ou ; trapezoidal except for slightly concave sinuate margin, and rounded PL corners ; posterior 
margin slightly convex but flattened medially. AL and PL setae stoutly and densely branched, the 
proximal 6-8 branches, blade-like; the apical ones more appressed, appearing scale-like. AM seta 
proximally with rows of 6-7 short, stout barbules, apically the setules more slender. AM seta inserted on 
a non-punctate depressed area with lateral margins marked by a longitudinal, somewhat angled ridge ; 
rest of scutum punctate except for another smooth area behind each SB. Sensillae about 45-50(4 long; 
shaft proximally smooth; apical half with about 14 relatively conspicuous branches. 
STUD. INST. MED. RES. 
