TROMBICULA FROM BORNEO 
49 
Standard Measurements in Microns, T. baluensis n. sp. 
AW 
PW 
SB 
ASB 
PSB 
A-P 
AL 
AM 
PL 
DS 
PW 
Cox II 
PW 
SD 
PW 
ASB 
PW 
A-P 
Holotype ... 
64 
78 
35 
27 
15 
30 
55 
53 
62 
59 
78/60 
= 1.3 
78/42 
= 1.9 
78/27 
= 2.9 
78/30 
=2.6 
Paratypes (8) Mean 
63 
79 
35 
28 
15 
30 
54 
54 
61 
59 
1.28 
1.85 
2.8 
2.6 
Range ( + or — ) 
2 
3 
I 
2 
i 
2 
3 
4 
3 
4 
0.05 
0.07 
O.I 
0.05 
Body Setae: Dorsal setae about 40-555 including lateromarginals, frequently arranged 2. 10. 10. 10. 
8... the rest confused; anteriormost about 55-61^ long. Dorsal setae of characteristic appearance, with 
a stout shaft and 5-8 conspicuous widely separated blade-like barbs, with a few closely appressed scale-like 
barbules along length of shaft. True ventral setae ca. 25-30;^ long with many slender branches and no 
blade-like processes, about 22 in number, plus 4-8 caudal setae (postanal), somewhat longer with fewer 
branches and mostly with a few blade-like processes. Legs: Sensory setae as in akamushi group, i.e.. Leg I 
with 2 genuala and a microgenuala ; with 2 tibialae (one stout) and a microtibiala ; with a blunt striate 
tarsala (spur), a microtarsala posterior and slightly distal to it, a well- developed subterminala and 
parasubterminala one-half its length, a pretarsala. Leg II with a genuala; 2 tibialae, one of which is stout 
and subapical; a tarsala and anterior microtarsala, a well-developed subterminala. Leg III with a genuala 
and a tibiala. 
Type Material. — Holotype (C.O.R.U. 20946), and 2 paratypes ex Crocidura , a true shrew, Mt. 
Kinabalu, Tenompok, elev. 4500 ft., 17. vii. 1951. ( R . Traub and J. R. Audy ). Nine paratypes, ibid , 
but ex Hylomys suillus the lesser gymnura or short-tailed shrew, and one paratype ex Dremomys everetti , a 
ground squirrel. Four additional paratypes ex 2 Hylomys and one from Rattus sp., 1 oc. cit ., v. 1952 ( J . R. 
Audy). Holotype (U.S.N.M. 2014); paratypes distributed as for T. flagellifera n. sp. 
Trombicula (Leptotrombidium) pipellae n. sp. (figs. 46-53, pp. 62, 63). 
Diagnosis of Larva . — Close to T. (L). longiseta Womersley, 1952 and T. deliensis , Walch, 
1922. Separable from all other known species of ‘ akamushi group’ except longiseta by the 
relatively enormous length of the posterolateral setae (over 1207., cf. maximum of 85 in deliensis 
— the PLs are nearly 2\ times as long as ASB + PSB) and by the very long dorsal setae (a mean 
of over 85^, cf. a maximum of 67 in deliensis) ; AM also much longer (an average of 87, not 
56 as in T. longiseta ), but AL nevertheless of same approximate length (56^). Distinguishable 
from longiseta (and deliensis and allies) by the relative and actual greater width of the scutum — 
i.e., PW with a mean of 92 instead of 72 as in longiseta. A-P about 37, not 26^. The palpal 
claw is 2-pronged as in T. longiseta ; it is trifurcate in T. deliensis. ( See fig. 43, p. 60) 
Description of Larva. — Body: About 446 x 28o[j. in moderately engorged holotype; very large in fully 
engorged individuals (1120x800^.). Eyes 2+2, on ocular plate, distinct, subovate, but posterior eye 
usually slightly longer and narrower than anterior. Gnathosome: Chelicers each with a small subapical 
tooth or notch but lacking a more proximal denticle. Palpal formula N/N/BNN. Galeal seta B 
(sparsely). Palpal claw apparently bifurcate. Scutum: Subrectangular ; somewhat less than twice as 
broad as long; anterior margin very shallowly biconcave; lateral margins somewhat concave; posterior 
margin slightly sinuous (reflecting the SBs). Sensillary bases inserted at level slightly anterior to PLs. 
Sensillae nude except for subapical short branches; about 65-7014 long. AL setae with short barbs; 
somewhat longer than scutum (ASB-)-PSB = SD, Scutal Depth). PL setae about twice length of AL, or 
even longer; similar in structure. Body Setae: About 26-28 dorsal setae arranged 2. 6(8). 6. 6... the rest 
irregular; anteriormost about 85-9210. long; with many short inconspicuous barbs. True ventral setae 
about 50[o., subpectinate, about 14 VS, plus 6-10 caudal setae (CS) ca. 7010.. Sensory setae as in T. 
baluensis n. sp. and in “ akamushi group”. 
Standard Measurements in Microns, T. pipellae n. sp. 
AW 
PW 
SB 
ASB 
PSB 
A-P 
AL 
AM 
PL 
DS 
PW 
Cox II 
PW 
SD 
PW 
ASB 
PW 
A-P 
Holotype ... 
78 
9 i 
33 
40 
14 
37 
59 
85 
123 
87 
91/78 
= 1.16 
, 91/54 
= 1.7 
91/40 
=2.3 
91/37 
= 2.46 
Paratypes (19) Mean 
80 
93 
32 
39 
15 
38 
61 
89 
125 
91 
1. 15 
i -7 
2.38 
2-44 
Range (+ or — ; ... 
3 
3 
2 
2 
2 
2 
3 
5 
8 
7 
0.05 
n 05 
0.07 
0.05 
Type Material. — Holotype (B9013-1) and 21 paratypes ex Prionodon linsang gracilis Horsfield, a civet 
(Viverridae), Mt. Kinabalu, Tenompok, elev. 4,500 ft., 17. vii. 1951 ( R . Traub and J. R. Audy). Sixteen 
other paratypes as follows : ex Callosciurus notatus , ibid , 18. vii; 6 ex Helictis everetti , a ferret-badger, 
ibid, 8, vii; 1 ex Rattus alticola , ibid, 16. vii; 4 ex Rattus mulleri, Jesselton, 8. vii. 1951 (D. H. Johnson 
MALA YA, No. 26, 1953 
