TROMBICULA FROM BORNEO 
5i 
Trombicula (Leptotrombidium) tarsala n. sp. (figs. 62-69, PP- 66, 67). 
Diagnosis of Larva. — Near T. deliensis but separated at once by the enlarged striate tarsala 
(spur) on Leg I, being about as long as the tarsus is broad at this point, instead of scarcely 
half so. The tarsala II correspondingly enlarged, and in about the same proportion to breadth 
of tarsus. 
Description of Larva. — Body: About 351x24514 in fairly engorged holotype. Eyes 2+2 on ocular 
plate, the anterior eye slightly the larger. Gnathosome: Chelicers each with a tricuspid cap. Palpal 
formula N/N/BNN. Palpal claw perhaps 3-pronged, but not determinable in specimens extant. Galeal 
seta B. Scutum almost two-thirds long as broad; posterior margin somewhat convex, evenly so; anterior 
margin essentially slightly concave but medially minutely sinuate; lateral margins sloping gently mesad 
towards AL with short “ shoulders ” at insertion of PLs. AM slightly longer than PLs which in turn are 
half-again as long as ALs; the scutal bristles barbed, the barbs rather appressed in AM and PL. Sensillae 
nude for basal two-thirds, then lightly branched. Bases of sensillae inserted slightly anterior to level of 
PLs. Body Setae: Dorsal setae about 26 in number arranged 2. 6. 6. 4. 4. 2: similar to PLs. True 
ventral setae 26-3054 plumose, numbering 14-16, plus about 6 caudal setae, ca. 40-520. resembling DS but 
at times more plumose. Legs: Sensory setae as in T. baluensis but with tarsala of Leg I relatively very 
large, submarginal, slightly curved, about subequal to breadth of tarsus at level of base of tarsala; tarsala 
of Leg II as in I but more median. 
Standard Measurements in Microns, T. tarsala n. sp. 
AW 
PW 
SB 
ASB 
PSB 
A-P 
AL 
AM 
PL 
DS 
PW 
Cox II 
PW 
SD 
PW 
ASB 
PW 
A-P 
Holotype ... 
59 
70 
28 
28 
11 
26 
42 
68 
66 
62 
70/59 
= 1.22 
70/39 
= 1.8 
70/28 
= 2.5 
70/26 
=2.7 
Paratype 
59 
66 
26 
28 
12 
27 
45 
69 
69 
63 
70/58 
= 1.20 
70/40 
= 1-75 
70/28 
= 2.5 
II 0 
Type Material. — Holotype (C.O.R.U. 21314) ex Rattus mulleri , Forest Reserve, mile 5 Stapok Road, 
Kuching, Sarawak, 30. viii. 1952; and one Paratype (No. 21288), same host species and locality, 31. viii. 
52 ( Tom Harrisson). 
Trombicula (Leptotrombidium) micula n. sp. (figs. 87-94, pp. 72, 73). 
Diagnosis of Larva. — Close to T. deliensis but with a scutum which is actually smaller 
and proportionately less deep (PW with a mean of 63 microns, SD = 3i), the ratio PW/SD=2, 
not 1.7-1.8, and PW/A-P is 3.1, not 2.2-5; SB nevertheless similar in both species (28-3054); 
posterior margin of scutum rather evenly convex, instead of being slightly convex or quite 
straight and immediately behind PLs ; sensillae bases inserted at the level of PLs or slightly 
behind, not somewhat anterior to PLs. 
Description of Larva. — Body: About 201 X 13754 in partially engorged holotype. Eyes 2+2 on ocular 
plate; posterior eye somewhat the smaller. Gnathosome: Chelicers each with the usual subapical notch. 
Palpal formula N/N/B(b)NN but dorsal tibial seta only slightly frayed, not truly barbed. Palpal claw 
3 -pronged, Galeal seta B. Scutum: About twice as broad as long; subrectangular but for rather arcuate 
posterior margin, anterior and lateral margins fairly straight except for typical bulge at level of insertion 
of ALs or PLs. PL slightly longer than AM which is somewhat longer than ALs: these setae with short 
cilia or barbs, but longest on AL. Sensillae with proximal half nude; apical half lightly branched. 
Body Setae: About 28 dorsal setae usually arranged 2. 8. 6. (7). 6. 4. 2.; similar to PLs. True ventral 
setae ca. 2ijx, sparsely plumed, numbering about 16, plus 6-8 cadual setae, postanal, 26a., resembling DS. 
Legs: sensory setae as in baluensis n. sp. and others of “akamushi group”. 
Standard Measurements in Microns, T. micula n. sp. 
AW 
PW 
SB 
ASB 
PSB 
A-P 
AL 
AM 
PL 
DS 
PW 
Cox 11 
PW 
SD 
PW 
ASB 
PW 
A-P 
Holotype ... 
58 
62 
30 
20 
11 
20 
27 
35 
40 
38 
62/50 
= 1.24 
62/31 
= 2.0 
62/20 
= 3-1 
62/20 
= 3-1 
Paratypes (8) Mean 
56 
62 
29 
19 
12 
20 
28 
35 
42 
39 
1.27 
2.0 
3-1 
3-1 
Range 
3 
4 
I 
2 
1 
2 
2 
3 
4 
3 
0.05 
0.06 
0.09 
0.02 
Type Material: Holotype and 7 paratypes ex Nannosciurus melanotis, a pigmy squirrel, Sarawak, 
Kuching, Stapok Forest Reserve, xi. 1951. ( Tom Harrisson ). Two paratypes ex Callosciurus notatus, 
ibid. Holotype (U.S.N.M. 2017) in U.S. National Museum. Paratypes in that institution and in 
collections of Colonial Office Medical Research Unit and of R. Traub. 
MALA YA , No. 26 , 1953 
