98 
TRAUB & AUDY 
Description of Larva. — Engorged specimen 364x298^. Gnathosome: as in G. birella n. sp., scutum 
also similar but lateral margins more evenly rounded, not as angulate anteriorly and lacking a median 
constriction; more finely punctate. Sensillae absent in specimens extant. Body Setae: Bases of dorsal 
Standard Measurements in Microns, G. arona n. sp. 
Slide 
AW 
PW 
PPW 
SB 
ASB 
PSB 
Holotype ... 
37 
64 
62 
37 
21 
85 
Paratypes (15) 
Mean 
35 
60 
57 
35 
22 
82 
Range ( + or 
-) 
2 
4 
5 
3 
2 
4 
A-P 
A-PP 
PP 
AL 
PL 
DS 
PW 
Cox II 
PW 
SD 
34 
69 
37 
29 
26 
26 
62/52 
= 1.15 
62/106 
= 0.59 
34 
66 
35 
30 
26 
27 
59/50 
= x.18 
59/104 
= 0.57 
2 
4 
2 
2 
2 
3 
— 
— 
and ventral setae somewhat enlarged, prominent. Dorsal setae relatively long and thin, about 28 or 30 in 
number ; 3 or 4 of these singly in an irregular line midway between lateral margins of scutum and sides of 
body. Ventral setae about 36 in number, about 15^ in length. Anal plate (fig. 37) ovate, about 50 X 36(4 
nearly as well sclerotized as scutum ; the valvular sclerite around anal aperture well demarcated. 
Type Material. — Holotype (U.S.N.M. 2069; C.O.R.U. 29214). and two paratypes ex Rattus mulleri 
(Jentinck), Malaya, Selangor, Ulu Langat, 5.viii.i952 (J. R. Audy ), in U.S. National Museum. Two 
paratypes, with same data, in British Museum of Natural History, and 5 ibid , in collection of Colonial 
Office Medical Research Unit, Kuala Lumpur. Two paratypes in collection of senior author, ibid, but 
ex Rattus whiteheadi (Thomas), Selangor, Kepong. Twelve additional paratypes with same data as 
holotype, distributed among various acarological collections. 
Comment . — The nymph of this species has been bred by Messrs. Nadchatram and Lee 
Fatt-Hing of the Colonial Office Unit. It is a typical Gahrliepiine and the anal plate appears to 
have no homologue in the nymphs. The nymph is to be described later (Womersley and 
Audy, in MS). 
Gahrliepia (Schongastiella) kalrata n. sp. (figs. 44-52). 
Diagnosis of Larva . — Near G. (S.) ceylonica Womersley, 1952, in the scutum being markedly 
and narrowly ligulate posterior to PL and in possessing multisetose coxae III. Separable 
from that species in that the scutum is proportionately much larger, AW ranges from 47-56^, 
not 31 and PW about 47-58^, not 28. Proportionately large size of scutum reflected in ratio 
of PW/Coxa-II, which is over 0.84; in ceylonica it is less than 0.72. With over 80 ventral 
setae instead of approximately 60. Coxae III typically bearing 5 setae each and only rarely 4. 
Description of Larva. — Approximately 680 by 480 [i in engorged specimen. Eye apparently single, 
at level between AL and SB. Gnathosome: with femoral setae slightly barbed or plumed (other palpal 
setae nude as in liota n. sp. and related forms). Scutum: markedly narrowed beyond PL so that scutum is 
flask-shaped; apex rapidly becoming subacuminate beyond PPL. Pseudo-posterolateral bristles (PPL) 
often inserted at different levels, so that bases are in an oblique line, not horizontal. Body Setae: Dorsal 
Standard Measurements in Microns, G. kalrata n. sp. 
Slide 
AW 
PW 
PPW 
SB 
ASB 
PSB 
Holotype ... 
56 
58 
19 
47 
28 
98 
Paratypes (8) 
Mean 
50 
53 
17 
43 
26 
89 
Range 
( + or — ) ... 
4 
6 
3 
4 
3 
12 
A-P 
A-PP 
PP 
AL 
PL 
DS 
PW 
Cox II 
PW 
SD 
47 
IOI 
25 
29 
36 
3 i 
58/60 
= 0.96 
58/126 
= 0.45 
46 
94 
25 
29 
36 
34 
0.98 
o .47 
5 
1 1 
8 
3 
2 
2 
0.08 
0.02 
setae as scutal setae but slightly smaller; about 70 in number, arranged in rows beginning 4. 8. (10) 10 (12) 
and therefore quite variable. Ventral setae about 85 in number, anteriormost about 2ip. in length, 
posterior ones about 28. Coxae III usually with 5 setae; at times with 5 on 1 and 4 on other, or 4-4; the 
most dorsal of these inserted at upper lateral angle of coxa, at times inapparent. 
Type Material. — Holotype (U.S.N.M. 2071) ex Rattus rattoides ssp. (turkestanicus ?), Kashmir, 
Sonmarg, viii.1949 (S.L. Kalra ). Six paratypes, ibid, or ex ‘ Mouse ’, Kashmir, Baltal, ibid, deposited in 
collections of the U.S. National Museum, the British Museum, and of the senior author. 
Comment . — The specific name was suggested by that of the collector, Lt-Col. S. L. Kalra, 
who has contributed much to our knowledge of scrub typhus. We are indebted to Dr. Audy 
for allowing us to study and describe this material, which was sent to him by Kalra. 
STUD. INST. MED. RES. 
