1 1 8 
H. WOMERSLEY 
In the general conformity of the scutum this genus might have come in the subfamily 
Apoloniinae of the Leeuwenhoekiidae, particularly in having leg I 7-segmented. It differs 
from the other genera of the subfamily in that only Leg I is 7-segmented. In the palpal claws 
it resembles Womersleyia Radford, and in the single tarsal claws it closely resembles 
to Neotrombidium Leonardi ( Monunguis Wharton). 
Nymphs of this species have however been bred by Messrs. Lee Fatt-Hing and Aman bin 
Mahmood of the Colonial Office Research Unit, and these are clearly related to Trombella Berl. 
in the subfamily Trombellinae. A description of the nymph, together with a review of the 
whole subfamily, is to be published elsewhere. 
The genus is named after Dr. J. R. Audy, leader of the British Colonial Office Research 
Unit in Malaya. 
Type Audy ana thompsoni n. sp. 
Audyana thompsoni n. sp. (figs. 15 A-F). 
Description of Larvae. — Size (engorged) to 650 jj., width to 450^. Shape oval. Legs: I 285(1. long, 
II 235j 1, III 26o,u. Scutum as figured with paired AM, AL and PL seate and 2 sensillae; anterior margin 
concave, apparently without any median process, but with a deep hyaline band extending the whole width 
of scutum ; AM and AL setae modified to short claviform bodies, and AL not at the antero-lateral corners 
but placed close to, lateral to and in line with sensillae bases j PL long and very narrowly lanceolate; 
sensillae longer and distinctly lanceolate. Eyes apparently absent. Palpi stout, tibia with two short 
straight claws, maxillae and femora with short claviform setae; genu with a long nude seta; tibia with 3 
nude setae ; tarsus conical over-reaching tibial claws, with two long straight apical spines and 4 or 5 straight 
nude setae. Chelicerae strongly curved, with a minute spical tricuspid cap, and two small subapically dorsal 
teeth. Dorsal seta tapering, finely but indistinctly ciliated, arranged 2.6.64.4.2 to 58^ long. Ventrally, 2 
fine ciliated setae on coxae I, a single claviform seta on coxae II and III, a pair of fine ciliated setae bet- 
ween coxae III, then 4.44.2 setae. Legs: I 7-segmented, II and III 6-segmented; all tarsi with only a 
single claw; no long nude seta on tarsi III. 
The 
Standard Data 
FOR 8 SPECIMENS 
ARE : 
Mean. 
Standard 
Theoretical 
Observed 
Coeff. of 
Deviation. 
Range. 
Range. 
Variation. 
AWSf ... 
71.05+1.05 .. 
. 2.97+0.74 . 
.. 62.15-79.95 
... 70.0-78.4 
4.1 
AWt 
57.05+0.74 .. 
2.08+0.52 
.. 50.8-63.3 
... 36.0-61.6 
3-6 
PW 
93. 4+0.94 .. 
2.65 + 0.66 . 
.. 85.4-101.4 
... 89.6-98.0 
2.8 
SB 
33- 6+0.53 .. 
1.50+0.37 
.. 29.1-38.1 
... 30.8-36.4 
4.4 
ASB 
30.45 + 0.35 .. 
. 0.99+0.25 
.. 27.5-33.4 
... 28.0-30.8 
3.2 
PSB 
16.8 
No variation recorded 
SD 
47.25 + 0.35 .. 
0.99+0.25 . 
.. 44-3-50.2 
... 44.8-47.6 
2.1 
(A-P)St ... 
37.1 + 0.70 
1.98+0.49 
.. 31.2-43.0 
... 33-6-39-2 
5-3 
AM 
11. 2 
No variation recorded 
AL 
11. 2 
No variation recorded 
PL 
84. 0+1.59 .. 
4.49+ 1. 12 
•• 70.5-97-5 
... 78.4-89.6 
5-3 
Sens. 
76. 3 + 2.10 .. 
4.2+1.48 
.. 63.7-88.9 
... 70.0-78.4 
5-5 
Loc. and Host. — Eight specimens (C.O.R.U. 4244-7) taken from the venter of the common giant 
black scorpion, Heterometrus longimanus (No. R 6049), Pahang Road, Ulu Gombak Forest Reserve, 
Selangor, Malaya, 14.iv.1949 ( J . R. Audy). Many more specimens have been collected since. 
Remarks. — This species is named for Lieut. -General Sir Treffry O. Thompson, K.c.s.i., 
C.B., c.b.e., k.h.p., d.m., d.p.h., formerly Director of Medical Services, Allied Land Forces, 
South East Asia, in recognition of his great encouragement of research on Scrub Typhus during 
World War II. 
Audyana thompsoni is reported to occur in small numbers, usually attached near the pecten, 
on nearly half the specimens of Heterometrus collected from forest in Selangor. As already 
noted, nymphs have been bred out, enabling the species to be placed in the Trombellinae. 
t In the above, AWS is the anterior width of the scutum, and AW is the width between the transposed AL setae/measured 
as defined for the Standard Data. Similarly (A-P)S is the distance between the antero-lateral corners of the scutum and the 
PL bases. 
STUD. INST. MED. RES. 
