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J. R. AUDY 
distinctive nymph resembles those of “ VN ” and “ BL” in shape (ASL/SB ratio about 
1.2 ?) and in the sensillae having sparse long branchlets and spiculate shaft; the long, solitary, 
paracristal setae are however peculiar. Mr. Womersley is kindly providing material for further 
study. The peculiar location (conjunctiva) of oculicola on the host is interesting in view of the 
occurrence of “ NS ” n. sp. on the intranasal mucosa. One of Vercammen’s species noted 
below was also intranasal. It must however be admitted that Vercammen has shown that a 
number of unrelated species favour this peculiar habitat. 
The relationships of at least four other species to this group require study: E. andrei (Jadin 
and Verc., 1952:622), E. crocidura (Law. 1949) “ DBI-NB ” n. sp. Jadin and Verc. (1953 in 
press) and E. nausheraensis (Worn. 1952:202) A specimen of the third was very kindly sent to 
the writer by Vercammen- Grand jean of the Laboratoire Medicale Province du Kivu, Bukavu 
(Belgian Congo). It has a small scutum similar in shape to that of E. andrei , the 3-pronged 
palpal claws are characteristically curved terminally, the chelicers have the single large tooth, 
and coxa III is multisetose. E. crocidura is closely similar. This species may link E. andrei 
to the group under discussion. Womersley’s species nausheraensis , of which the writer has 
kindly been lent several specimens, has a somewhat different scutum (larger, PW more nearly 
equal to AW, SBs relatively wide apart) but the posterior border is concave and the scutum 
generally resembles that of andrei (in this, Womersley’s figure 52C is slightly inaccurate). 
There are only 2 humeral setae but coxa III is multisetose. The palpal claw is 2-pronged and 
not particularly short or incurved, resembling that of “ NS ” n. sp. from Borneo. Vercammen’s 
species are in a number of features intermediate between nausheraensis and oculicola , and a good 
deal of study will be necessary to clarify these interesting relationships. T. praomyia Rad. 
from Africa and T. jubbulporensis Worn, have a peculiar scutum like those of E. andrei and 
E. nausheraensis : they have been described as Trombicula in the absence of sensillae and their 
true relationships are unknown. 
Old World species, relationships sub-judice. — Africa: ? andrei, ? crocidura, “ DBI-NB ” Jad. & Verc. 
Pdutoiti. Asia: nausheraensis, oculicola. 
Genus Riedlinia Oudemans, 1914 
Type. — Riedlinia coeca Ouds. 1914:88, Ent. Ber. Amst. 4, 84-49. From a bat, hills near Bombay, 
India. 
Diagnosis. — Monotypic: larva with spatulate empodium, claws stronger than usual; eyes absent; 
scutum roughly pentagonal, with posterior angle, sensillae unknown. Nymphs not known. 
Remarks. — Womersley describes this species as having globose sensillae but Fuller 
(1952:200) in a redescription states that the nature of these organs is unknown. Kalra is 
attempting to collect topotypic material (personal communication). There is a lapsus calamae in 
Wharton and Fuller where the locality is given as Malaya. 
Genus Endotrombicula Ewing, 1931 
Type. — Endotrombicula penetrans Ewing, 1931 :i6, Proc. U.S. Nat. Mus., 80, 1-19. From amphibian. 
East Africa. 
Diagnosis. — Trombiculines with expanded sensillae, chelicers with large recurved teeth increasing in 
size proximally; apparently confined to amphibians. 
Subgenus Endotrombicula, Wharton and Fuller, 1952 
Type. — Endotrombicula penetrans Ewing, 1931 :i6. From amphibian, East Africa. 
Diagnosis. — Endotrombicula whose larvae have a single tarsala I. 
Old World species (Africa). — madagascariensis ; penetrans ; pillersi. 
Subgenus Phrynacarus Lawrence, 1949, Wharton and Fuller, 1952 
Type. — Phrynacarus fitzsimonsi Lawrence, 1949:463, Ann. Natal. Mus., 11, 405-486. From 
amphibian. South Africa. 
Diagnosis. — Monotypic; Endotrombicula whose larva has several (4) tarsalae I. 
STUD. INST. MED. RES. 
