i68 
J. R. AUDY 
5. In addition to the accepted genera and subgenera, over 30 species groups are listed, at 
least three of which may deserve recognition as subgenera (viz. the Euschongastia indica , 
E. lacunosa , and Doloisia oculicola groups). 
6. The Trombiculinae are divided into two generic groups, related respectively to 
Trombicula and to Euschongastia. The latter group has many affinities with the Gahrliepiinae 
(Walchiinae). The genera Walchiella (here treated as a subgenus of Euschongastia , close to the 
lacunosa-g roup) and Pseudoschongastia are considered to belong properly to the Euschongastia 
complex and not to the Gahrliepiines. 
7. The genus Trombicula sensu lato is revised and the subgenus Trombicula is based on the 
larval and nymphal characters of a new, undescribed bat-chigger which appears to be congeneric, 
if not identical, with the genotype T. minor. Evidence is offered to show that the postlarval 
stages of T. minor lack eyes. The subgenus Leptotrombidium is provisionally revised sensu 
lato ; over 60 species are at present allocated to it. 
8. Fonsecia is restored to generic rank and it is considered that the T. ilesi group, which 
does not have modified scutal setae, may prove to be congeneric with it. 
9. The genus Euschongastia sensu lato is provisionally considered to comprise the subgenera 
Euschongastia , Walchiella , and Helenicula n. subgen., together with the indica-g roup and the 
lacunosa-gvoup which appear to be at least of subgeneric status but the limits of which cannot 
yet be defined. The lacunosa-g roup may possibly be identified with Walchiella , while the 
indica-g roup has close affinities with Doloisia sensu lato. 
10. The genus Doloisia is treated sensu lato, comprising ( a ) the subgenus Doloisia sensu 
stricto, (b) a new subgenus (to be published), the larvae of which have PL setae off the scutum 
and the nymphs of which are very distinctive, resembling those of Guntherana and the 
Gahrliepiines in the shape of the scutum, and (c) an oculicola-complcx which requires much 
further study. The larvae of Doloisia appear to have affinities with members of the E. indica 
group, and this requires further study. 
11. The Gahrliepiinae (Walchiinae) are treated sensu stricto because they are a very 
distinctive group in both larvae and nymphs. The broad genus Gahriiepia is considered to be 
the temporary repository of two or more groups which will later be given generic status. It is 
considered that the existing subgenera are based on false distinctions and that the subfamily 
needs drastic revision. 
12. The Leeuwenhoekiinae and Apoloniinae are hardly represented in the Indo-Malaysian 
collections, with the exception of Whartonia on bats. These subfamilies are briefly considered. 
Larval characters of the various genera are tabulated: Womersley’s new genera, based on 
chiggers from arthropods and provisionally placed by him in the Apoloniinae, are included in the 
tabulation. 
References 
Audy, J. R. (1947). Scrub typhus as a study in ecology. Nature , Load ., 159, 295-296. 
Audy, J. R. (1952). Trombiculid mites infesting bats in Malaya, with descriptions of three new species. 
Bull. Raffles Mus., 24 , 133-159* 
Audy, J. R. (1953). Malaysian parasites. Ill — A summary review of collections of trombiculid mites 
in the Asiatic-Pacific area. Stud. Inst. med. Res., Malaya, 26 , 29-44. 
Audy, J. R. (1954). Malaysian parasites. New trombiculid species of taxonomic interest. Stud. Inst, 
med. Res., Malaya (in preparation). 
Audy, J. R., Thomas, H. M. and Harrison, J. L. (1953). A collection of trombiculid mites from Manipur 
and Lower Burma, 1945-1946. J. Zool. Soc. India, 5, 20-40. 
Baker, J. A. and Wharton, G. W. (1952). An introduction to acarology. Macmillan, New York. 
STUD. INST. MED. RES. 
