3 . 966 ] 
Reichardt — Eucheilini 
9 
This small Neotropical tribe includes only two genera of small, 
Lebiini-like Carabidae. Their systematic position has been uncertain 
for a long time. Even though Eucheila , the type-genus, was described 
as early as 1829 (in Lebiini), the true relations to Helluonini were 
only recognized in 1883, when Bates erected the subfamily Eucheilinae 
to incorporate Eucheila and Inna (the latter also described in Lebiini, 
and at first considered as related to Eucheila by Bates), and placed it 
in the vicinity of the Helluonini. 
In 1929 Liebke described the subfamily Periglossinae for a new 
Central-American genus, Periglossium. From his description and 
illustrations of this beetle, it is evident that Periglossium is a synonym 
of Inna, and consequently the name Periglossinae has to be suppressed. 
The characters which link the Eucheilini to the Helluonini are 
the strangely modified mouthparts (Figs. 1-8). In spite of sim- 
ilarities, the Eucheilini are undoubtedly a distinct tribe, easily dis- 
tinguished from the Lebiini by the completely different mouthparts 
and from the Helluonini by their general Lebiini-habitus, as well as 
by the antennae which are pubescent from the 4th segment on in 
Eucheilini (pubescent from base in Helluonini). The aedeagus of 
the Eucheilini was unknown up to now. I was able to dissect one 
male of Eucheila strandi (Liebke) and one of Inna boyeri (Solier) 
(see Figs. 11 and 12). The two aedeagi are very similar, and this 
fact strengthens the supposed relation between the two genera. The 
left paramere of the aedeagus is reduced, but still present, being 
somewhat lobate in the two species. The genitalia of Neotropical 
Helluonini are also unknown, so that no comparison can be made 
now. 
Geographic distribution: The tribe is typically Neotropical, ex- 
tending from Argentina to the southern United States (Texas). No 
species has yet been reported from the Antilles. 
Key to genera 
1. Labrum convex, covering apex of mandibles, with short setae 
only on lateral margins (Fig. 8) ; lateral margins of pronotum 
smooth, not crenulated; pronotum with basal setae only; tarsal 
claws pectinate Eucheila Dejean 
Labrum flat, not covering apex of mandibles, with 4 long setae 
on anterior margin, 2 longer ones and a series of short ones 
laterally (Fig. 3); lateral margins of pronotum crenulated; 
pronotum with basal and latero-median setae; tarsal claws 
simple Inna Putzeys 
