Psyche 
[March 
5§ 
highly variable; cross veins numerous and regularly arranged over 
the areas of Rs, M and CuA; an irregular network in costal area 
and between CuA and CuP and the anal veins. Wing membrane 
with fine microtrichia between veins; prominent setae or other cuti- 
cular derivatives developed to variable degrees on most of the veins 
of wing, including many cross veins. 
Hind wing unknown and bodjr structure unknown, except for part 
of one leg. 
This family has not been given any diagnosis previously, except 
by generic assignment. In the preceding account I have attempted to 
bring together venational characteristics of the fore wing present in 
several Palaeozoic genera which are apparently closely related to 
Strephocladus and which I am placing in the family. However, the 
strephocladids will probably turn out to be an extensive group and 
its diagnosis will undoubtedly need modification as other genera be- 
come known. 
Strephocladus was established in the Order Palaeodictyoptera by 
Scudder (1885) for a species ( subtilis Kliver) which was collected 
in Upper Carboniferous strata of Saarbrucken and which had orig- 
inally been placed by Kliver in the blattod genus Petroblattina. It 
has subsequently been placed in the order Protoblattoidea, Incertae 
Sedis, by Handlirsch (1908, 1921); in the order Protorthoptera, 
family Oedischiidae, by Waterlot (1934) and Guthorl (1936); in 
a new order Strephocladodea, family (new) Strephocladidae, by Mar- 
tynov (1938); in the order Paraplecoptera, family Strephocladidae 
by Sharov ( 1961) ; and in the Protorthoptera, family Strephocladidae, 
by Kukalova (1965), who added another genus, Spargoptilon (L. 
Permian, Moravia), to the family. From my study of the type speci- 
men of Strephocladus subtilis and of the several species from Elmo 
Text-figure 1. Strephocladus subtilis Kliver. Original drawing of fore 
wing, based on holctype, No. D/164, Bergingenieurschule, Saarbrucken. 
