92 
Psyche 
[June 
and by fine granulation; costal expansion unknown; wings relatively 
broad and short, apical part prolonged and narrow; anterior margin 
strongly convex, posterior margin slightly concave apically; branches 
of main veins nearly parallel, distinct; Sc sending off several parallel 
branches, almost leaching apex; R sending off a series of long branches 
towards Sc ; Rs hardly distinguishable from the other branches of R ; 
M and CuA with a variable number of irregular branches; CuP 
long, branched, slightly concave; anal veins about 5 in number, some 
directed anteriorly; cross veins numerous, regular. 
Relationship. Protocoleus Tillyard, 1924, differs from the related 
new genus Phyllelytron in the more distinct venation, in the more 
regularly and parallel-arranged veins and cross veins, and in the 
regularly arranged tubercles, distinct on the whole wing surface. 
From Austrelytron , n. gen., it differs in the more richly branched main 
veins, the presence of many cross veins and absence of regular series 
of pointed tubercles on the veins. 
Stratigraphic occurrence: Upper Permian of Australia (N.S.W.). 
Type-species: Protocoleus initchelli Tillyard, 1924 
Protocoleus mitchelli Tillyard 
Protocoleus initchelli Tillyard, 1924, Proc. Linn. Soc., N.S.W. 49(4) :432; 
pi. 46, fig. 3. 
Fore wing: about 27-29 mm. long and 7-9.3 mm. broad, sutural 
margin broader in the proximal half, narrowing abruptly in the distal 
half; tubercles densely arranged, often bordering veins in regular 
rows; fore wing very narrow in the apical third; anterior margin 
very convex, in the distal third slightly concave; branches of R dis- 
tinct, very oblique and regular; cross veins mostly straight. 
Discussion. Since Tillyard had at his disposal only the reverse of 
Protocoleus for study, he considered the surface of the elytron to be 
covered by pits. Actually, the tubercles and interspaces between them 
are so much alike that the reverse and obverse sculpturing does not 
show much difference. The fine granulation, on the other hand, ap- 
pears in the reverse half of the fossil as small, dense pits. The 
granulation, which is extraordinarily fine in Protocoleus ■, was not 
mentioned in Tillyard’s original description. It is very probable that 
both tubercles and granulation were present on both dorsal and ventral 
surfaces of the tegmina. 
Holotype: specimen (reverse), figured by Tillyard (1924) on 
plate 46, fig. 3 ; collected in Upper Permian beds of Belmont, N.S.W. 
(Not seen). 
