1966] 
Kukalova — Protelytroptera 
99 
In the Tillyard collection of the British Museum (N.H.) in Lon- 
don, I found 4 specimens, corresponding to this description. One of 
them (No. In 45518) is designated by the label, attached to the 
fossil and in Tillyard’s hand-writing, as “Permophilus?” . For that 
reason, there is little doubt that the fossils belong to the family Per- 
mophilidae. They are clearly protelytropterous, showing the costal 
expansion (interpreted by Tillyard as an ailula), distinct sutural 
margin, the setae in patches, and, in spite of its reduction, a protely- 
tropterous pattern of the venation. Laurentiaux, it should be noted 
(1953, P- 475), referred the Permophilidae to a new order Para- 
coleoptera, which is not distinct from the Protelytroptera. 
The wing surface of Permophilidae (dense granulation and tu- 
bercles), as well as the form of the wings, resembles that in the 
family Protocoleidae. It is not impossible that they might have 
originated from a common ancestor. 
Geographical occurrence of family: Upper Permian of Australia 
(N.S.W.). 
Genera included: Permophilus Tillyard, 1924; Elytrathrix, new 
genus. 
Genus Permophilus Tillyard 
Permophilus Tillyard, 1924, Proc. Linn. Soc., N.S.W. 49:434; Jeannel, 1949, 
Traite de zool. 9:64; Laurentiaux, 1953, Traite de paleont. 3:476. 
Fore wing : tegminous, slightly convex, darkly pigmented ; sutural 
margin relatively narrow; wing surface covered by dense granulation 
and isolated, indistinct tubercles; costal expansion projecting; apical 
part narrowed and prolonged; main veins strongly developed in the 
basal third, but absent completely in the distal part of the wing. 
Relationship. Not having at my disposal the type for study, I am 
not able to decide definitely the generic position of Specimens No. In 
46014 and In 45518. According to the original diagnosis given by 
Tillyard, they differ from the holotype of P. pincombei only in the 
smaller size. For this reason, 1 refer them to the same genus, but 
they might also be considered as representing a separate genus. Speci- 
men No. In 45517 (fragment) corresponds to about the size given 
for the type of Permophilus, but it is too poor for any conclusion. 
Specimen No. In 45503, which I refer to a new genus Elytrathr’ix , 
has very distinct, large tubercles concentrated at the basal part of 
the fore wing, which would presumably have been mentioned in Till- 
yard’s description of Permophilus pincombei if they had been present. 
Stratigraphic occurrence: Upper Permian of Australia (N.S.W.) 
Type-species: Permophilus pincombei Tillyard, 1924 
