A REVIEW OF HALOCORYZA ALLUAUD, 
WITH NOTES ON ITS RELATIONSHIP 
TO SCHIZ 0 GENIUS PUTZEYS 
(COLEOPTERA: CARABIDAE). 
By Donald R. Whitehead 
Department of Entomology, University of Alberta 
INTRODUCTION 
My work on a revision of Nearctic species of the scaritine genus 
Schizogenius Putzeys has brought to light two species, one of them 
new, that properly belong to the otherwise African Halocoryza 
Alluaud. The major purposes of this paper are to help clarify the 
taxonomy of Halocoryza , to describe the American species, and to 
contrast the genus against Schizogenius. Halocoryza is poorly known 
and apparently very rare in collections. Despite this limitation, how- 
ever, I think some interesting zoogeographic and phylogenetic points 
emerge. These are considered briefly, following the taxonomic 
treatment. 
Halocoryza and Schizogenius belong to the “tribe” Clivinini 
{sensu Jeannel, 1946), within which they may be recognized by 
having a bi- or trituberculate clypeal margin (the antero-lateral 
angles angulate but not tuberculate) , along with four to six pairs 
of nearly parallel longitudinal ridges between the eyes. In the closely 
related Lophocoryza Alluaud, the clypeal margin is quadrituberculate 
and the paired frontal carinae are reduced, the outer pairs short and 
oblique. At least one other genus, Coryza Putzeys, should probably 
be placed in the same group ; but it is only distantly related, and 
lacks paired ridges on the frons. 
Ball’s (i960) key to North American scaritine genera, as modified 
by the substitution of couplet 3(1) below, can be used to distinguish 
Schizogenius and Halocoryza from all other Scaritini of the world. 
3(1). Frons with four to six pairs of longitudinally directed 
carinae between eyes, and clypeus bi- or trituberculate 
along anterior margin 3A. 
Frons with two deep grooves, or with shallow, transverse 
or oblique grooves, but without a series of parallel carinae 
(North America) ; or, if frons with four or more pairs 
of nearly parallel longitudinal carinae between eyes, clypeus 
without tubercles along anterior margin (World) 4. 
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