1966] 
Chapin — Coccinellidae 
279 
Figure 1. Brachyacantha drntipcs (Fabricius). A. Aedeagus (without 
sipho) ; B. Sipho ; C. Ferna^ eenitalia, showing receptaculum seminis, sperm 
duct, infundibulum, and apex of bursa. 
cantha Chevrolat, Cyra Mulsant, Hinda Mulsant, and probably 
Cleothera Mulsant. These genera are characterized in the males 
by parameres attached to the tegmen in the normal manner for 
Coccinellidae, that is, attached to the basal ring at the base of the 
median lobe and without special “roots” which pass through or almost 
through the ring; in the females by the presence of a complicated 
infundibular structure and by the possession of a normally formed, 
more or less sausage-shaped, receptaculum seminis. The infundibulum 
in these genera consists of three or four arms, two or three of which 
lie in the wall of the bursa copulatrix and act as supporting struts. 
The remaining arm stands free, and the sperm duct joins the bursa 
at the junction of these arms. The sperm duct is relatively short. 
The receptaculum lacks well-developed nodulus or ramus. These 
genera also agree in having eleven-segmented antennae and toothed 
tarsal claws. 
Hyperaspis Redtenbacher, Hyperaspidius Crotch, Helesius Casey, 
Thalassa Mulsant, Corystes Mulsant, and Oxynychus Leconte and 
probably Diazonema Weise, Tiphysa Mulsant, and Menoscelis Muls- 
ant, are included in the second division. These genera are character- 
ized in the males by parameres which appear to be outgrowths of 
sclerotized but unpigmented structures or “roots,” within or ex- 
tending beyond the basal ring; in the females by the possession of a 
compound receptaculum seminis, and by the absence of an infundibu- 
lum. Among these genera one finds species with nine, ten, or eleven 
antennal segments and with toothed or toothless claws. 
